Bark and Ambrosia Beetles of

SYNONYMY

• Pityophthorus laetus Wood 1976. Great Basin Nat. 36(3): 358

DISTRIBUTION.

Numbers in parentheses after each geographic unit are the number of distinct collection events in the database for that unit. For exotic species generally only countries are listed for localities outside the New World. For further information on published sources of distribution, check the REFERENCES section.

• Central America (5)
  Costa Rica (1): Heredia (1); Guatemala (1): unknown (1); Panamá (3): Darién (2), Panamá (1);
• South America (37)
  Bolivia (9): Bení (1), Cochabamba (1), Santa Cruz (7); Brazil (5): Bahia (1), Espirito Santo (1), Pará (1), Rôndonia (2); Colombia (3): Putomayo (2), unknown (1); Ecuador (1): Pastaza (1); Fr. Guiana (1): unknown (1); Perú (11): Cusco (1), Huanuco (2), Huánuco (1), Junin (2), Loreto (2), Madre de Dios (3); Suriname (1): unknown (1); Venezuela (6): Aragua (2), Barinas (1), Mérida (1), Miranda (2);

HOSTS

Numbers in parentheses after each host family, genus, or collection method are the number of distinct collection events in the database for that host or method.

• No host information (26);
• Leguminosae (1): Lonchocarpus (1): margaritensis (1);
• Malvaceae (1): Theobroma (1): cacao (1);
• Melastomataceae (2): (2): no species data (2);
• trap (12) light (12): no details (2), mercury vapor + UV (3), ultraviolet (7);

REFERENCES

The following are important recent monographs, catalogs, and supplements to catalogs that refer to this species. The specific page on which the reference is made is shown in pink at the end of the reference. In the case of Wood (1982) and Wood & Bright (1992) clicking on the reference page will link to a digital version of the work in question.

• Bright, D.E. 1981. A taxonomic monograph of the genus Pityophthorus Eichhoff in North and Central America (Coleoptera: Scolytidae). Entomol. Soc. Can. Mem. 118:1-378. [103]. (data capture complete)
• Wood, S.L. 1982. The bark and ambrosia beetles of North and Central America (Coleoptera: Scolytidae), a taxonomic monograph. Great Basin Nat. Mem. 6:1-1356. [1081]. (data capture complete)
• Wood, S.L., Bright,D.E. 1992. A catalog of Scolytidae and Platypodidae (Coleoptera), Part 2. Taxonomic Index. Great Basin Nat. Mem. 13:1-1553 (vol. A, B). [1005]

The following are references from which host and distribution data have been input into the database. If one of the above monographs or catalogs also appears in this list, it means that most relevant collection event data have been included.

1. Bright, D.E. 1981. A taxonomic monograph of the genus Pityophthorus Eichhoff in North and Central America (Coleoptera: Scolytidae). Entomol. Soc. Can. Mem. 118: 1-378.
2. Wood, S.L. 1982. The bark and ambrosia beetles of North and Central America (Coleoptera: Scolytidae), a taxonomic monograph. Great Basin Nat. Mem. 6: 1-1356.
3. Wood, S.L.; Bright, D.E. 1992. A catalog of Scolytidae and Platypodidae (Coleoptera), Part 2. Taxonomic Index (Volumes A,B). Great Basin Nat. Mem. 13: 1-1553.

Maps automatically open at the center of the plotted points and the scale is set to encompass all map points. Maps can be resized (scale bar at upper left) and the center moved (place cursor over map and drag) to see other parts of the distribution of the species. Clicking on a map marker will pull up collection event data and a literature citation if present. If any errors are found, please refer to the "series code" which is a unique identifier for a database record in any communications). Coordinates have not been entered for all collection records. Localities outside the New World are not plotted, even though they are listed in the distribution summary and in the table of records.

No.	Collection data	Publication
Central America
1->	Costa Rica, Heredia, Volcan Poas; 19/XI/1963; Wood, S.L.	[USNM (1: holotype, female) (Bright, D.E. 1981: 103) (Wood, S.L. 1982: 1081) (Wood, S.L.; Bright, D.E. 1992: 1005)] [USNM (2: paratype(s), female) (Bright, D.E. 1981: 103)]
2->	Guatemala, unknown, Guatemala (country)	[unknown (1) (Wood, S.L.; Bright, D.E. 1992: 765)]
3->	Panamá, Darién, Cana Field Station; 29/VI/2009; Sekerka, L.; Gowin, E.; Windsor, D.; trap: light	[BMNH (1) (TH Atkinson-2016, unpublished)]
4->	Panamá, Darién, Cerro Pirre; 3/VI/2017; Riley, E.G.; trap: light ultraviolet	[TAMU (1) (TH Atkinson-2019, unpublished)]
5->	Panamá, Panamá, Ft. Clayton, Albrook Forest; 1967; Hutton, R.; trap: light ultraviolet	[UAAM (2) (TH Atkinson-2007, unpublished)]
South America
6->	Bolivia, Bení, Rurrenabaque, 20 km E; 13/X/2004; Wappes, J.E.; Morris, R.	[FSCA (1) (TH Atkinson-2016, unpublished)]
7->	Bolivia, Cochabamba, Villa Tunari; 10/IX/2012; Skelley, P.E.; Wappes, J.E.; trap: light mercury vapor + UV	[FSCA (3) (TH Atkinson-2016, unpublished)]
8->	Bolivia, Santa Cruz, Buena Vista, 3.7 km SSE, Hotel Flora & Fauna; 22/XI/2013; Wappes, J.E.; Skilman	[FSCA (1) (TH Atkinson-2016, unpublished)]
9->	Bolivia, Santa Cruz, Buena Vista, 3.7 km SSE, Hotel Flora & Fauna; 5/XI/2001; Thomas, M.C.; trap: light ultraviolet	[FSCA (3) (TH Atkinson-2016, unpublished)]
10->	Bolivia, Santa Cruz, Buena Vista, 3.7 km SSE, Hotel Flora & Fauna; 14/X/2000; Thomas, M.C.; trap: light ultraviolet	[FSCA (2) (TH Atkinson-2016, unpublished)]
11->	Bolivia, Santa Cruz, Buena Vista, 3.7 km SSE, Hotel Flora & Fauna; 19/X/2004; Wappes, J.E.; Morris, R.	[FSCA (2) (TH Atkinson-2016, unpublished)]
12->	Bolivia, Santa Cruz, Buena Vista, 3.7 km SSE, Hotel Flora & Fauna; 15/XI/2001; Thomas, M.C.; Dozier, B.K.; trap: light ultraviolet	[FSCA (1) (TH Atkinson-2016, unpublished)] [UTIC (2 , female) (TH Atkinson-2016, unpublished)]
13->	Bolivia, Santa Cruz, Cafetal; 5/VIII/1990; Parillo, P.; Bettella, P.	[FMNH (1) (TH Atkinson-2022, unpublished)]
14->	Bolivia, Santa Cruz, Potrerillos de Guendá, 40 km NW Santa Cruz; 12/X/2007; Cline, A.R.; Wappes, J.; trap: light mercury vapor + UV	[CSCA (1) (TH Atkinson-2021, unpublished)]
15->	Brazil, Bahia, Ilheus; 11/III/1981; Kaston	[unknown (1) (Wood, S.L. 2007: 458)]
16->	Brazil, Espirito Santo, Aracruz; 18/XI/1992	[MEFEIS (1) (Wood, S.L. 2007: 458)]
17->	Brazil, Pará, Belém; XI/1977; Ohashi, O.	[MEFEIS (1) (Wood, S.L. 2007: 458)]
18->	Brazil, Rôndonia, Ariquemes, 62 km SW; 28/X/1996; Vulinec, K.; trap: light mercury vapor + UV	[FSCA (2) (TH Atkinson-2007, unpublished)]
19->	Brazil, Rôndonia, Ariquemes, 62 km SW; 23/VIII/1996; Schmitz, U.; trap: light ultraviolet	[FSCA (1) (TH Atkinson-2007, unpublished)]
20->	Colombia, Putomayo, Santa Rosa; 4/X/1970; Malkin, B.; Burchard, P.	[FMNH (2) (TH Atkinson-2010, unpublished) (DE Bright-1984, unpublished)]
21->	Colombia, Putomayo, Santa Rosa de Sucumbios; 25/VIII/1971; Malkin, B.	[FMNH (1) (DE Bright-1984, unpublished) (TH Atkinson-2013, unpublished)]
22->	Colombia, unknown, Colombia (country)	[ZUMH (1: holotype, female) (Wood, S.L.; Bright, D.E. 1992: 765) (Wood, S.L. 2007: 458)]
23->	Ecuador, Pastaza, Ashuara, Rio Macuma, 10 km Rio Morona; 7/VII/1971; Malkin, B.	[FMNH (2) (TH Atkinson-2010, unpublished) (DE Bright-1984, unpublished)]
24->	Fr. Guiana, unknown, Fr. Guiana (country)	[unknown (1) (Wood, S.L.; Bright, D.E. 1992: 765)]
25->	Perú, Cusco, Marcapata	[unknown (1) (Smith, S.M.; Beaver, R.A.; Cognato, A.I. 2017: 92)]
26->	Perú, Huanuco, Leonpampa; 11/XII/1937; Woytkowski, F.	[unknown (1) (Wood, S.L. 2007: 458) (Smith, S.M.; Beaver, R.A.; Cognato, A.I. 2017: 92)]
27->	Perú, Huanuco, Leonpampa; 23/IX/1935; Woytkowski, F.	[unknown (1) (Wood, S.L. 2007: 458)]
28->	Perú, Huánuco, Tingo Maria; 18/IX/1954; Schlinger, E.I.; Ross, E.S.	[unknown (1) (Wood, S.L. 2007: 458) (Smith, S.M.; Beaver, R.A.; Cognato, A.I. 2017: 92)]
29->	Perú, Junin, Canan Eden, 8 km NNE Puerto Ocopa, north bank of Perene Riv; Petrov, A.V.	[AVP (6) (Smith, S.M.; Beaver, R.A.; Cognato, A.I. 2017: 92)]
30->	Perú, Junin, Río Venado, 15 km NW Satipo; Petrov, A.V.	[AVP (3)
31->	Perú, Loreto, 40 km NE Iquitos on Amazon Riv., Explorama Lodge; 22/VIII/1992; Castner, J.; Skelley, P.E.; trap: light ultraviolet	[FSCA (3) (TH Atkinson-2007, unpublished) (Smith, S.M.; Beaver, R.A.; Cognato, A.I. 2017: 92)]
32->	Perú, Loreto, Tambo Pirana, Río Cochiquinas; 30/VI/1978; Hespenheide, H.A.	[TAMU (1) (TH Atkinson-2013, unpublished)]
33->	Perú, Madre de Dios, Cocha Cashu Biological Station, Manu National Park; 18/X/2000; Brooks, R.	[SEMC (2) (TH Atkinson-2020, unpublished)]
34->	Perú, Madre de Dios, Los Amigos Biol. Station; Smith, S.M.; Hulcr, J.	[MSUC (1) (TH Atkinson-2007, unpublished)]
35->	Perú, Madre de Dios, Tambopata Res.; 12/I/1987; Wheeler, G.C.; trap: light	[CUIC (1) (TH Atkinson-2010, unpublished)]
36->	Suriname, unknown, Suriname (country)	[unknown (1) (Wood, S.L.; Bright, D.E. 1992: 765) (Wood, S.L. 2007: 458)]
37->	Venezuela, Aragua, Colonia Tovar; 4/V/1970; Wood, S.L.; Melastomataceae	[USNM (1: holotype, male) (Wood, S.L. 2007: 761)] [USNM (22: paratype(s)) (Wood, S.L. 2007: 761) (TH Atkinson-2017, unpublished)] [USNM (1: allotype, female) (Wood, S.L. 2007: 761) (TH Atkinson-2017, unpublished)]
38->	Venezuela, Aragua, Colonia Tovar; 4/V/1970; Wood, S.L.; Melastomataceae	[USNM (1: allotype, female) (Wood, S.L. 2007: 761)] [USNM (11: paratype(s)) (Wood, S.L. 2007: 761)]
39->	Venezuela, Barinas, El Canton, 40 km E; 8/III/1970; Wood, S.L.; Leguminosae: Lonchocarpus margaritensis	[USNM (1) (Wood, S.L. 2007: 458)]
40->	Venezuela, Mérida, Mérida; 28/II/1970; Wood, S.L.	[USNM (21: paratype(s)) (Wood, S.L. 2007: 761)]
41->	Venezuela, Miranda, Cacagua; 1971; Saunders, J.L.; Malvaceae: Theobroma cacao	[unknown (1) (Wood, S.L. 2007: 458)]
42->	Venezuela, Miranda, El Laurel Experimental Farm, 12 km SW Caracas; 1/V/1970; Wood, S.L.	[USNM (1: paratype(s)) (Wood, S.L. 2007: 761)]