Bark and Ambrosia Beetles of

SYNONYMY

• Pityoborus rubentis Wood 1958. Great Basin Nat. 18: 51

DISTRIBUTION.

Numbers in parentheses after each geographic unit are the number of distinct collection events in the database for that unit. For exotic species generally only countries are listed for localities outside the New World. For further information on published sources of distribution, check the REFERENCES section.

Distribution Comments.  In ranges of western and central Mexico.

• Asia (15)
  China (9): no state / prov. data (2), Fujian (2), Hong Kong (1), Taiwan (3), Yunnan (1); India (1): Assam (1); Nepal (1): no state / prov. data (1); Thailand (1): Chiang Mai (1); Vietnam (3): no state / prov. data (3);
• Central America (26)
  (3): no state / prov. data (3); Costa Rica (13): Cartago (5), Heredia (2), Limón (2), Puntarenas (1), San José (3); Guatemala (1): Amatitlan (1); Honduras (3): Atlántida (1), Olancho (2); Panamá (6): Chiriquí (3), Coclé (1), Panamá (2);
• Eurasia (12)
  France (8): no state / prov. data (4), Auvergne-Rhône-Alpes (4); Japan (2): no state / prov. data (1), Gansho (1); Korea (2): Jeju-do (1), Jeonlanam-do (1);
• North America (24)
  México (16): Chiapas (3), Chihuahua (1), Durango (2), Jalisco (2), México (1), Michoacán (1), Oaxaca (1), Puebla (1), Sonora (1), Veracruz (3); United States (8): Alabama (1), California (1), Florida (3), South Carolina (3);
• South America (2)
  Perú (1): Madre de Dios (1); Venezuela (1): unknown (1);

HOSTS

Numbers in parentheses after each host family, genus, or collection method are the number of distinct collection events in the database for that host or method.

Host Comments.  In shaded out branches of living trees.

• No host information (34);
• Adoxaceae (1): Sambucus (1): sp. (1);
• Anacardiaceae (1): Spondias (1): purpurea (1);
• Burseraceae (1): Protium (1): sp. (1);
• Dipterocarpaceae (1): Vatica (1): lanceaefolia (1);
• Fagaceae (6): (1): no species data (1); Castanea (1): crenata (1); Castanopsis (2): sp. (2); Cyclobalanopsis (1): glauca (1); Pasania (1): sp. (1);
• Leguminosae (2): Pentaclethra (2): macroloba (2);
• Malvaceae (1): Theobroma (1): cacao (1);
• Melastomataceae (2): Conostegia (2): oerstediana (2);
• Pinaceae (8): Abies (1): firma (1); Pinus (7): no species data (1), engelmannii (1), oocarpa (1), sp. (4);
• trap (22) flight intercept (1): no details (1); light (3): no details (1), mercury vapor + UV (1), ultraviolet (1); Lindgren Funnel (7): no details (3), alpha pinene + ethanol (1), ethanol (1), ips-lure (1), UHR ethanol (1); soda bottle (2): no details (1), ethanol (1); sticky trap (1): no details (1); window trap (8): no details (8);

REFERENCES

The following are important recent monographs, catalogs, and supplements to catalogs that refer to this species. The specific page on which the reference is made is shown in pink at the end of the reference. In the case of Wood (1982) and Wood & Bright (1992) clicking on the reference page will link to a digital version of the work in question.

• Wood, S.L. 1982. The bark and ambrosia beetles of North and Central America (Coleoptera: Scolytidae), a taxonomic monograph. Great Basin Nat. Mem. 6:1-1356. [1150]. (data capture complete)
• Wood, S.L., Bright,D.E. 1992. A catalog of Scolytidae and Platypodidae (Coleoptera), Part 2. Taxonomic Index. Great Basin Nat. Mem. 13:1-1553 (vol. A, B). [968]

The following are references from which host and distribution data have been input into the database. If one of the above monographs or catalogs also appears in this list, it means that most relevant collection event data have been included.

1. . . .
2. Atkinson, T.H. 2019. Escarabajos descortezadores y ambrosiales (Coleoptera: Curculionidae: Scolytinae, Platypodinae) de Sonora, México (Bark and Ambrosia Beetles (Coleoptera: Curculionidae: Scolytinae, Platypodinae) of Sonora, México). Dugesiana 26(1): 41-49.
3. Wood, S.L. 1982. The bark and ambrosia beetles of North and Central America (Coleoptera: Scolytidae), a taxonomic monograph. Great Basin Nat. Mem. 6: 1-1356.

Maps automatically open at the center of the plotted points and the scale is set to encompass all map points. Maps can be resized (scale bar at upper left) and the center moved (place cursor over map and drag) to see other parts of the distribution of the species. Clicking on a map marker will pull up collection event data and a literature citation if present. If any errors are found, please refer to the "series code" which is a unique identifier for a database record in any communications). Coordinates have not been entered for all collection records. Localities outside the New World are not plotted, even though they are listed in the distribution summary and in the table of records.

No.	Collection data	Publication
Asia
1->	China, no state / prov. data; 27/IV/2010; Duarte, A.; Muhammad, A.; trap: soda bottle	[UGCA (2) (Hoebeke, E.R.; Rabaglia, R.J.; Knížek, M.; Weaver, J.S. 2018: 248)]
2->	China, no state / prov. data, Beijing; 15/IV/1980; Yu, P.	[USNM (1) (Smith, S.M.; Cognato, A.I. 2022: 9)]
3->	China, Fujian, Chong’an, Guidun; 7/V/1978; Fagaceae: Cyclobalanopsis glauca	[USNM (2) (Smith, S.M.; Cognato, A.I. 2022: 9)]
4->	China, Fujian, Kuatun	[ZFMK (1: holotype, female) (Wood, S.L.; Bright, D.E. 1992: 740) (Smith, S.M.; Cognato, A.I. 2022: 9)]
5->	China, Hong Kong, Tai Po Kau; VI/2017; Skelton, J.; Fagaceae: Castanopsis sp.	[MSUC (2) (Smith, S.M.; Cognato, A.I. 2022: 9)]
6->	China, Taiwan, Fushan; V/2009; trap: sticky trap	[RJRC (1) (Hoebeke, E.R.; Rabaglia, R.J.; Knížek, M.; Weaver, J.S. 2018: 248)]
7->	China, Taiwan, Fushan; III/2015; Hulcr, J.; Fagaceae: Pasania sp.	[UFFE (1) (Smith, S.M.; Cognato, A.I. 2022: 9)]
8->	China, Taiwan, Taichung, Heping Dist.; 29/IV/2015; Lin, C.S.	[MSUC (1) (Smith, S.M.; Cognato, A.I. 2022: 9)]
9->	China, Yunnan, Xishuangbanna, Sanchahe Nat. Res; 29/V/2008; Cognato, A.I.	[MSUC (4) (Smith, S.M.; Cognato, A.I. 2022: 9)]
10->	India, Assam, Nakachari, Sibsigar; 17/V/1921; Beeson, C.F.C.; Dipterocarpaceae: Vatica lanceaefolia	[FRIDD (1: neotype, female) (Smith, S.M.; Beaver, R.A.; Singh, S.; Cognato, A.I. 2018: 139) (Smith, S.M.; Beaver, R.A.; Cognato, A.I. 2020: 186) (Smith, S.M.; Cognato, A.I. 2022: 9)]
11->	Nepal, no state / prov. data, Nepal	[unknown (1) (Wood, S.L.; Bright, D.E. 1992: 740)]
12->	Thailand, Chiang Mai, Doi Pui; Fagaceae: Castanopsis sp.	[unknown (1) (Beaver, R.A.; Sittichaya, W.; Liu, L.Y. 2014: 38)]
13->	Vietnam, no state / prov. data, Cao Bang; Cognato, A.I.; Smith, S.M.; Pham	[MSUC (4) (Smith, S.M.; Cognato, A.I. 2022: 9)]
14->	Vietnam, no state / prov. data, Lao Cai, Hoang Lien NP; 17/V/2019; Smith, S.M.; Cognato, A.I.	[MSUC (1) (Smith, S.M.; Cognato, A.I. 2022: 9)]
15->	Vietnam, no state / prov. data, Thua Thien-Hue, Bach Ma NP; 14/II/2017; Cognato, A.I.; Hoang, T.A.; Fagaceae	[MSUC (2) (Smith, S.M.; Cognato, A.I. 2022: 9)]
Central America
16->	, no state / prov. data; Melastomataceae: Conostegia oerstediana	[unknown (0) (Wood, S.L.; Bright, D.E. 1992: 1072)]
17->	, no state / prov. data; Anacardiaceae: Spondias purpurea	[unknown (0) (Wood, S.L.; Bright, D.E. 1992: 1072)]
18->	, no state / prov. data; Malvaceae: Theobroma cacao	[unknown (0) (Wood, S.L.; Bright, D.E. 1992: 1072) (Blandford, W.F.H. 1905: 259)]
19->	Costa Rica, Cartago, Cartago; Wood, S.L.	[USNM (1) (Wood, S.L. 1982: 1309)]
20->	Costa Rica, Cartago, Peralta; Wood, S.L.	[USNM (1) (Wood, S.L. 1982: 1309)]
21->	Costa Rica, Cartago, Tapanti; 17/VIII/1963; Wood, S.L.	[USNM (1: holotype, female) (Wood, S.L. 1982: 1309)]
22->	Costa Rica, Cartago, Tapanti; 2/VII/1963; Wood, S.L.; Melastomataceae: Conostegia oerstediana	[USNM (1) (Wood, S.L. 1982: 1309) (TH Atkinson-2024, unpublished)]
23->	Costa Rica, Cartago, Turrialba; 13/III/1965; Duckworth, S.S.; Duckworth, W.D.	[USNM (1 , female) (TH Atkinson-2024, unpublished)]
24->	Costa Rica, Heredia, La Selva Biol. Sta., 3 km S Pto. Viejo; 5/IV/1984; Hespenheide, H.A.; Leguminosae: Pentaclethra macroloba	[HAHC (2) (TH Atkinson-1989, unpublished)]
25->	Costa Rica, Heredia, La Selva Biol. Sta., 3 km S Pto. Viejo; 5/IV/1984; Hespenheide, H.A.; Leguminosae: Pentaclethra macroloba	[TAMU (1) (TH Atkinson-1989, unpublished)]
26->	Costa Rica, Limón, Beverley; Wood, S.L.	[USNM (1) (Wood, S.L. 1982: 1309)]
27->	Costa Rica, Limón, Río Reventazon, Hamburgfarm; 2/II/1934; Nevermann, F.	[FMNH (1) (TH Atkinson-2024, unpublished)]
28->	Costa Rica, Puntarenas, Volcan; Wood, S.L.	[USNM (1) (Wood, S.L. 1982: 1309)]
29->	Costa Rica, San José, Escasú; Wood, S.L.	[USNM (1) (Wood, S.L. 1982: 1309)]
30->	Costa Rica, San José, San José; Wood, S.L.	[USNM (1) (Wood, S.L. 1982: 1309)]
31->	Costa Rica, San José, Santa Ana; Wood, S.L.	[USNM (1) (Wood, S.L. 1982: 1309)]
32->	Guatemala, Amatitlan, Tacatón; 2/IX/1995; MacVean, C.; trap: light ultraviolet	[CAS (1) (TH Atkinson-2007, unpublished)]
33->	Honduras, Atlántida, Parque Nal. Pico Bonito, El Portillo; 29/VI/2001; Turnbow, R.H.	[RHTC (1) (TH Atkinson-2010, unpublished)]
34->	Honduras, Olancho, Montaña del Malacate; 11/VI/2003; Turnbow, R.H.; trap: light mercury vapor + UV	[RHTC (1) (TH Atkinson-2010, unpublished)]
35->	Honduras, Olancho, Parque Nal. La Muralla; 24/V/1995; Wappes, J.E.	[FSCA (1) (TH Atkinson-2024, unpublished)]
36->	Panamá, Chiriquí, Bugaba; Wood, S.L.	[BMNH (1) (Blandford, W.F.H. 1905: 255)]
37->	Panamá, Chiriquí, El Hato del Volcán, 8 mi (13 km) S; 7/I/1964; Wood, S.L.	[USNM (1) (Blandford, W.F.H. 1905: 1309)]
38->	Panamá, Chiriquí, Volcán de Chiriqui; Champion, G.C.	[BMNH (1: holotype, female) (Wood, S.L. 1982: 1233)]
39->	Panamá, Coclé, Cerro Campana; 26/VI/1966; Wood, S.L.	[USNM (1) (Wood, S.L. 1982: 1309)]
40->	Panamá, Panamá, Camino del Oleoducto; 2/II/1999; Turnbow, R.H.	[RHTC (2) (TH Atkinson-2010, unpublished)]
41->	Panamá, Panamá, Isla Barro Colorado; 18/II/1971; Smythe, N.; trap: light	[TAMU (1) (TH Atkinson-2024, unpublished)]
Eurasia
42->	France, no state / prov. data, Crolles (38), ENS du Marais de Montfor; 2019; Dodelin, B.; trap: window trap	[unknown (1) (Dodelin, B.; Saurat, R. 2021: 0)]
43->	France, no state / prov. data, Engins (38), Réserve Biologique,; 2019; Dodelin, B.; trap: window trap	[unknown (1) (Dodelin, B.; Saurat, R. 2021: 0)]
44->	France, no state / prov. data, La Balme-Les-Grottes (38), ENS des Coteaux de Saint-Roch; 2019; Dodelin, B.; trap: window trap	[unknown (1) (Dodelin, B.; Saurat, R. 2021: 0)]
45->	France, no state / prov. data, Saint-Pierre-de-Chartreuse (38); 23/VI/2014; Janssen, P.; trap: window trap	[unknown (1) (Dodelin, B.; Saurat, R. 2021: 0)]
46->	France, Auvergne-Rhône-Alpes, Brison-Saint-Innocent (73); 2018; Dodelin, B.; trap: window trap	[unknown (1) (Dodelin, B.; Saurat, R. 2021: 0)]
47->	France, Auvergne-Rhône-Alpes, La-Motte-Servolex (73); 28/VI/2013; Dodelin, B.; trap: window trap	[unknown (2) (Dodelin, B.; Saurat, R. 2021: 0)]
48->	France, Auvergne-Rhône-Alpes, Pommier-la-Placette (38); 20/VI/2014; Janssen, P.; trap: window trap	[unknown (1) (Dodelin, B.; Saurat, R. 2021: 0) (Barnoiun, T,; Soldati, F.; Roques, A.; Faccoli, M.; Kirkendall, L.R.; Mouttet, R.; Daubree, J.B.; Noblecourt, T. 2020: 56)]
49->	France, Auvergne-Rhône-Alpes, Saint-Jean-de-Chevelu (73); 1/VIII/2017; Dodelin, B.; trap: window trap	[MSUC (1) (Dodelin, B.; Saurat, R. 2021: 0)]
50->	Japan, no state / prov. data, Tsukuba; 27/VIII/1980; Wood, S.L.; Pinaceae: Abies firma	[USNM (1) (Smith, S.M.; Cognato, A.I. 2022: 9)]
51->	Japan, Gansho, Shizuoka, Izu Peninsula; Fagaceae: Castanea crenata	[USNM (1: holotype, female) (Wood, S.L.; Bright, D.E. 1992: 740) (Smith, S.M.; Cognato, A.I. 2022: 9)]
52->	Korea, Jeju-do, province only	[unknown (1) (Park, S.; Smith, S.M.; Cognato, A.I.; Beaver, R.A. 2020: 213)]
53->	Korea, Jeonlanam-do, province only	[unknown (1) (Park, S.; Smith, S.M.; Cognato, A.I.; Beaver, R.A. 2020: 213)]
North America
54->	México, Chiapas, Metzabok; 1/VII/2021; Burgos S., A.	[UTIC (2) (TH Atkinson-2024, unpublished)]
55->	México, Chiapas, Metzabok, 3 km antes; 22/IX/2019; Burgos S., A.	[UTIC (4) (TH Atkinson-2021, unpublished)]
56->	México, Chiapas, Ocozocoautla, 13 mi N; 2/VIII/1969; Bright, D.E.	[CNC (1) (Bright, D.E. 1972: 1385)]
57->	México, Chihuahua, La Magdalena; 26/IV/1977; Furniss, M.M.	[unknown (1) (Wood, S.L. 1982: 1150)]
58->	México, Durango, Durango, 37 km W; 4/VI/1965; Wood, S.L.; Pinaceae: Pinus sp.	[USNM (1) (Wood, S.L. 1982: 1150)]
59->	México, Durango, El Salto, 3 mi (5 km) W; 7/VI/1965; Wood, S.L.; Pinaceae: Pinus sp.	[USNM (1) (Wood, S.L. 1982: 1150)]
60->	México, Jalisco, Cd. Guzmán; 14/IX/1978; Pinaceae: Pinus oocarpa	[INIF () (Atkinson, T.H.; Equihua M., A. 1988: 94)]
61->	México, Jalisco, Tlaquepaque; 14/XI/1940; Adoxaceae: Sambucus sp.	[USNM (1: holotype, male) (Bright, D.E. 1981: 58) (Wood, S.L. 1982: 1132) (TH Atkinson-2017, unpublished)] [USNM (165: paratype(s)) (Bright, D.E. 1981: 58)] [CNC (2: paratype(s)) (Bright, D.E. 1981: 58)]
62->	México, México, Texcoco; 5/II/1962; Coronado, R.; Pinaceae: Pinus sp.	[USNM (1) (Wood, S.L. 1982: 1150)]
63->	México, Michoacán, Volcán Paricutín, 6 mi (10 km) E; 19/VI/1965; Wood, S.L.; Pinaceae: Pinus	[USNM (1) (Wood, S.L. 1982: 1150)]
64->	México, Oaxaca, Uxpanapa; 26/V/1981; Equihua M., A.	[CEAM (2 , male) (Atkinson, T.H.; Equihua M., A. 1986: 422) (Atkinson, T.H.; Equihua M., A. 1986: 422) (TH Atkinson-2012, unpublished)]
65->	México, Puebla, Texmelucán, 14 mi (23 km) W; 14/VII/1953; Wood, S.L.; Pinaceae: Pinus sp.	[SEMC (1: holotype, female) (Wood, S.L. 1982: 1150)]
66->	México, Sonora, Mesa del Campanero; 11/III/2016; Atkinson, T.H.; Pinaceae: Pinus engelmannii	[UTIC (6) (Atkinson, T.H. 2019: 44)] [UAIC (4) (Atkinson, T.H. 2019: 44)] [TAMU (4) (Atkinson, T.H. 2019: 44)] [CEAM (3) (Atkinson, T.H. 2019: 44)]
67->	México, Veracruz, Laguna Escondida; 9/IX/2019; Ibarra J., L.A.; trap: Lindgren Funnel ethanol	[UTIC (1) (TH Atkinson-2019, unpublished)]
68->	México, Veracruz, Lake Catemaco; 20/VI/1969; Bright, D.E.	[CNC (1) (Bright, D.E. 1972: 1385) (Wood, S.L. 1982: 1233)] [USNM (2) (Wood, S.L. 1982: 1233) (TH Atkinson-2017, unpublished)]
69->	México, Veracruz, Lake Catemaco; 1/V/1969; Bright, D.E.	[CNC (1) (Bright, D.E. 1972: 1385)]
70->	United States, Alabama, Solon Dixon Center; 25/V/2017; Bares, H.M.; trap: Lindgren Funnel	[MEM (1) (Smith, S.M.; Cognato, A.I. 2022: 9)]
71->	United States, California, Felton; 17/VI/2019; trap: Lindgren Funnel UHR ethanol	[CDFA (1) (Smith, S.M.; Cognato, A.I. 2022: 9)]
72->	United States, Florida, Archer, 3.4 mi WNW; 7/VIII/2017; Schnepp, K.E.; trap: flight intercept	[UTIC (1) (TH Atkinson-2020, unpublished)] [KESC (1) (TH Atkinson-2020, unpublished)]
73->	United States, Florida, Florida Caverns SP; 26/IV/2017; Leahy; Danner; trap: Lindgren Funnel	[RJRC (1) (Smith, S.M.; Cognato, A.I. 2022: 9)]
74->	United States, Florida, Live Oak; 15/X/2020; Alford, B.; trap: Lindgren Funnel	[KESC (1) (TH Atkinson-2020, unpublished)]
75->	United States, South Carolina, North Charleston; 1/V/2012; Bishop, F.; trap: Lindgren Funnel alpha pinene + ethanol	[UGCA (1) (Hoebeke, E.R.; Rabaglia, R.J.; Knížek, M.; Weaver, J.S. 2018: 248)]
76->	United States, South Carolina, North Santee; 16/III/2012; Strickland, S.; trap: soda bottle ethanol	[UGCA (1) (Hoebeke, E.R.; Rabaglia, R.J.; Knížek, M.; Weaver, J.S. 2018: 248)]
77->	United States, South Carolina, Summerville; 14/V/2012; Bestwick, M.; trap: Lindgren Funnel ips-lure	[UGCA (1) (Hoebeke, E.R.; Rabaglia, R.J.; Knížek, M.; Weaver, J.S. 2018: 248)]
South America
78->	Perú, Madre de Dios, Los Amigos Biological Station; Smith, S.M.; Hulcr, J.; Burseraceae: Protium sp.	[MSUC (4) (Smith, S.M.; Petrov, A.V.; Cognato, A.I. 2017: 80)]
79->	Venezuela, unknown, Venezuela (country)	[unknown (1) (Wood, S.L.; Bright, D.E. 1992: 1056)]