Bark and Ambrosia Beetles of Americas

SYNONYMY

Taxonomic Comments.  Xyleborus intrusus is found from British Columbia to Gautemala, but is isolated from eastern populations of the same species by a large area in central Canada and the Lake states of the U.S. This species is very similar to X. pubescens Zimmermann from eastern North America, differing in minute differences in the denticles of the elytral declivity. Bright (2019:298) formally proposed the synonymy but this may be premature. In the meantime I believe that all records and names from eastern North America should be treated as pubescens.

• Xyleborus intrusus Blandford 1898. Biol. Cent. Amer. 4(6): 213
• Xyleborus fitchi Hopkins 1915. U.S. Dep. Agric. Tech. Bull. (99): 206
• Xyleborus howardi Hopkins 1915. U.S. Dep. Agric. Tech. Bull. (99): 65
• Xyleborus scopulorum Hopkins 1915. U.S. Dep. Agric. Tech. Bull. (99): 66

DISTRIBUTION.

Numbers in parentheses after each geographic unit are the number of distinct collection events in the database for that unit. For exotic species generally only countries are listed for localities outside the New World. For further information on published sources of distribution, check the REFERENCES section.

Distribution Comments.  Widely distributed in Western North America from southwestern Canada to CEntral America. A disjunct population is found in the northeastern U.S. and Appalachian regions. This eastern population may be synonymous with X. pubescens.

• Central America (6)
  El Salvador (1): San Salvador (1); Guatemala (1): Baja Verapaz (1); Honduras (4): Comayagua (1), El Paraíso (1), Francisco Morazán (2);
• Caribbean (1)
  Rep. Dominicana (1): unknown (1);
• North America (184)
  (4): no state / prov. data (4); Canada (1): British Columbia (1); México (32): Baja California (1), Baja California Sur (4), Chiapas (3), Distrito Federal (1), Durango (1), Guerrero (1), Hidalgo (1), Jalisco (2), México (4), Michoacán (1), Morelos (6), Oaxaca (3), Puebla (3), Tlaxcala (1); United States (147): Arizona (13), California (40), Colorado (20), District of Columbia (1), Florida (2), Georgia (3), Idaho (3), Louisiana (2), Maine (1), Maryland (2), Mississippi (4), Montana (2), Nebraska (1), Nevada (4), New Jersey (1), New Mexico (9), New York (2), North Carolina (7), Ohio (2), Oregon (4), Pennsylvania (9), Rhode Island (1), South Carolina (1), South Dakota (3), Utah (6), Virginia (1), West Virginia (3);

HOSTS

Numbers in parentheses after each host family, genus, or collection method are the number of distinct collection events in the database for that host or method.

• No host information (87);
• Pinaceae (18): Pinus (17): contorta (1), coulteri (1), duranguensis (1), jeffreyi (1), leiophylla (2), montezumae (2), oocarpa (3), patula (1), ponderosa (1), pseudostrobus (1), scopulorum (1), sp. (2); Pseudotsuga (1): menziesii (1);
• trap (86) (46): alpha pinene + ethanol (45), sugar bait (1); light (7): no details (1), mercury vapor + UV (3), ultraviolet (3); Lindgren Funnel (29): no details (1), alpha pinene + ethanol (14), ethanol (5), exotic ips lure (6), turpentine (1), turpentine + frontalin (2); pitfall (3): no details (3); window trap (1): no details (1);

REFERENCES

The following are important recent monographs, catalogs, and supplements to catalogs that refer to this species. The specific page on which the reference is made is shown in pink at the end of the reference. In the case of Wood (1982) and Wood & Bright (1992) clicking on the reference page will link to a digital version of the work in question.

• Wood, S.L. 1982. The bark and ambrosia beetles of North and Central America (Coleoptera: Scolytidae), a taxonomic monograph. Great Basin Nat. Mem. 6:1-1356. [839]. (data capture complete)
• Wood, S.L., Bright,D.E. 1992. A catalog of Scolytidae and Platypodidae (Coleoptera), Part 2. Taxonomic Index. Great Basin Nat. Mem. 13:1-1553 (vol. A, B). [745]

The following are references from which host and distribution data have been input into the database. If one of the above monographs or catalogs also appears in this list, it means that most relevant collection event data have been included.

1. . . .
2. Anonymous, 2010. Michigan State University. SCAN: Symbiota Collections of Artropods Network. http://symbiota4.acis.ufl.edu/scan/portal/collections/misc/collprofiles.php?collid=76: -.
3. Atkinson, T.H.; Saucedo C.; E, Martínez F.; E, Burgos S., A 1986. Coleópteros Scolytidae y Platypodidae asociados con las comunidades vegetales de clima templado y frío en el estado de Morelos. Acta Zool. Mex. 17: 1-58.
4. Bright, D.E. 1968. Review of the tribe Xyleborini in America north of Mexico (Coleoptera: Scolytidae). Can. Entomol. 100: 1288-1323.
5. Bright, D.E., Stark, R.W. 1973. The bark and ambrosia beetles of California. Coleoptera: Scolytidae and Platypodidae. Bull. Calif. Insect Surv. 16: 1-169.
6. EDRR 2011. USFS Early Detection Rapid Response Database. http://www.fs.fed.us/invasivespecies/earlydetection.shtml/.
7. EDRR 2014. USFS Early Detection Rapid Response Database. http://www.fs.fed.us/invasivespecies/earlydetection.shtml/.
8. Furniss, M.M., Johnson, J.B. 1987. List of Idaho Scolytidae (Coleoptera) and notes on new records. Great Basin Nat. 47: 757-762.
9. Hopkins, A.D. 1915. Classification of the Cryphalinae with descriptions of new genera and species. USDA Ent. Comm. Report (99): 1-75.
10. University of Arizona Insect Collection 2016. SCAN: Symbiota Collections of Arthropods Network. http://symbiota4.acis.ufl.edu/scan/portal/collections/misc/collprofiles.php?collid=11.
11. Weber, B.C., MacPherson, J.E. 1991. Seasonal Flight Patterns Of Scolytidae (Coleoptera) In Black-Walnut Plantations In North-Carolina And Illinois. Coleopterists Bull. 45(1): 45-56.
12. Wood, S.L. 1951. The Scolytidae of the Logan Canyon area in Utah and their host plants. Proc.Utah Acad.Sci.Arts Letters 26: 127-128.
13. Wood, S.L. 1982. The bark and ambrosia beetles of North and Central America (Coleoptera: Scolytidae), a taxonomic monograph. Great Basin Nat. Mem. 6: 1-1356.
14. Wood, S.L.; Bright, D.E. 1992. A catalog of Scolytidae and Platypodidae (Coleoptera), Part 2. Taxonomic Index (Volumes A,B). Great Basin Nat. Mem. 13: 1-1553.

Maps automatically open at the center of the plotted points and the scale is set to encompass all map points. Maps can be resized (scale bar at upper left) and the center moved (place cursor over map and drag) to see other parts of the distribution of the species. Clicking on a map marker will pull up collection event data and a literature citation if present. If any errors are found, please refer to the "series code" which is a unique identifier for a database record in any communications). Coordinates have not been entered for all collection records. Localities outside the New World are not plotted, even though they are listed in the distribution summary and in the table of records.

No.	Collection data	Publication
Central America
1->	El Salvador, San Salvador, El Boquerón; 10/VI/1959; Bechyné, J.	[IRSNB (1) (TH Atkinson-2013, unpublished)]
2->	Guatemala, Baja Verapaz, San Jerónimo	[BMNH (1: lectotype, female) (Wood, S.L. 1982: 839)]
3->	Honduras, Comayagua, Siguatepeque, 7 km E; 19/VII/1977; O'Brien, C.W.; O'Brien, L.B.; Marshall, G.B.	[CWOB () (Atkinson, T.H.; Equihua M., A. 1988: 102)]
4->	Honduras, El Paraíso, Yuscarán; 1/VI/2003; Turnbow, R.H.; trap: light mercury vapor + UV	[RHTC (2) (TH Atkinson-2008, unpublished)]
5->	Honduras, Francisco Morazán, El Zamorano	[unknown (1) (Wood, S.L. 1982: 839)]
6->	Honduras, Francisco Morazán, Talanga; 22/V/1996; Andrews, F.G.; Gilbert, A.J.; trap: light ultraviolet	[CSCA (2) (TH Atkinson-2014, unpublished)]
Caribbean
7->	Rep. Dominicana, unknown, Rep. Dominicana (country)	[unknown (1) (Wood, S.L.; Bright, D.E. 1992: 745)]
North America
8->	, no state / prov. data; Pinaceae: Pinus coulteri	[unknown (0) (Wood, S.L. 1982: 839)]
9->	, no state / prov. data; Pinaceae: Pinus jeffreyi	[unknown (0) (Wood, S.L. 1982: 839)]
10->	, no state / prov. data; Pinaceae: Pinus ponderosa	[unknown (0) (Wood, S.L. 1982: 839)]
11->	, no state / prov. data; Pinaceae: Pinus duranguensis	[unknown (0) (Wood, S.L. 1982: 839)]
12->	Canada, British Columbia, Aspen Grove	[unknown (1) (Wood, S.L. 1982: 839)]
13->	México, Baja California, Las Arrastras, 7 mi N.; 9/VI/1967; Sleeper, E.L.	[CAS (1) (TH Atkinson-2007, unpublished)]
14->	México, Baja California Sur, ramal a Los Naranjos, 24 km W; 28/VIII/1994; Turnbow, R.H.	[RHTC (1) (TH Atkinson-2008, unpublished)]
15->	México, Baja California Sur, , 2 mi E; 6/VII/1979; Andrews, F.G.; Hardy, A.R.; Giulani, D.	[CSCA (1) (TH Atkinson-2014, unpublished)]
16->	México, Baja California Sur, La Paz, 10 mi S; 26/VIII/1994; Turnbow, R.H.; trap: light mercury vapor + UV	[RHTC (2) (TH Atkinson-2008, unpublished)]
17->	México, Baja California Sur, Los Barriles, 3 km S; 27/VIII/1994; Turnbow, R.H.; trap: light mercury vapor + UV	[RHTC (1) (TH Atkinson-2008, unpublished)]
18->	México, Chiapas, Cintalapa, 20 km SW	[unknown (1) (Wood, S.L. 1982: 839)]
19->	México, Chiapas, San Cristobal de las Casas; 26/IX/1961	[CEAM () (Atkinson, T.H.; Equihua M., A. 1985: 238)]
20->	México, Chiapas, San Cristobal, 12.3 km W; 29/X/1986; Turnbow, R.H.	[RHTC (1) (TH Atkinson-1988, unpublished)]
21->	México, Distrito Federal, Xochimilco	[unknown (1) (Wood, S.L. 1982: 839)]
22->	México, Durango, El Salto	[unknown (1) (Wood, S.L. 1982: 839)]
23->	México, Guerrero, Chilapa; 23/II/1982; Atkinson, T.H.; Equihua M., A.; Pinaceae: Pinus oocarpa	[CEAM () (Atkinson, T.H.; Equihua M., A. 1985: 238)]
24->	México, Hidalgo, Tulancingo; 25/VI/1981; Pinaceae: Pinus patula	[UACC () (Atkinson, T.H.; Equihua M., A. 1988: 102)]
25->	México, Jalisco, Autlan, 6.1 m N; 30/VII/1996; Godwin, W.	[TAMU (1) (TH Atkinson-2003, unpublished)]
26->	México, Jalisco, Volcan Colima	[unknown (1) (Wood, S.L. 1982: 839)]
27->	México, México, Carr. Toluca-Morelia, km 54; 30/X/1980; Atkinson, T.H.; Equihua M., A.; Pinaceae: Pinus montezumae	[CEAM () (Atkinson, T.H.; Equihua M., A. 1985: 238)]
28->	México, México, Tlalmanalco	[unknown (1) (Wood, S.L. 1982: 839)]
29->	México, México, Toluca	[Unknown () (Wood, S.L. 1982: 839)]
30->	México, México, Toluca	[unknown (1) (Wood, S.L. 1982: 839)]
31->	México, Michoacán, Ario de Rosales; 30/X/1980; Atkinson, T.H.; Equihua M., A.; Pinaceae: Pinus pseudostrobus	[CEAM () (Atkinson, T.H.; Equihua M., A. 1985: 238)]
32->	México, Morelos, Carr. Xochimilco-Oaxtepec, km 56; Pinaceae: Pinus sp.	[CEAM () (Atkinson, T.H.; Saucedo C.; E, Martínez F.; E, Burgos S., A 1986: 35)]
33->	México, Morelos, Chamilpa; Pinaceae: Pinus montezumae	[CEAM () (Atkinson, T.H.; Saucedo C.; E, Martínez F.; E, Burgos S., A 1986: 35)]
34->	México, Morelos, Cuernavaca	[Unknown () (Wood, S.L. 1982: 839)]
35->	México, Morelos, Cuernavaca	[unknown (1) (Wood, S.L. 1982: 839)]
36->	México, Morelos, Cuernavaca, 13 km E. Hwy 160; 12/V/1983; O'Brien, C.W.; O'Brien, L.B.; Marshall, G.B.; trap: light	[FSCA (1) (TH Atkinson-2007, unpublished)]
37->	México, Morelos, El Guarda, 24 km S	[unknown (1) (Wood, S.L. 1982: 839)]
38->	México, Oaxaca, Benito Juárez; 5/II/1984; Equihua M., A.; Pinaceae: Pinus oocarpa	[CEAM () (Atkinson, T.H.; Equihua M., A. 1988: 102)]
39->	México, Oaxaca, Oaxaca	[Unknown () (Wood, S.L. 1982: 839)]
40->	México, Oaxaca, Sta. Ma. Chimalapa; 9/II/1984; Equihua M., A.; Pinaceae: Pinus oocarpa	[CEAM (10) (Atkinson, T.H.; Equihua M., A. 1988: 102) (TH Atkinson-2017, unpublished)] [DGSV (2) (TH Atkinson-2017, unpublished)]
41->	México, Puebla, Puebla; 17/I/1980; Atkinson, T.H.; Pinaceae: Pinus leiophylla	[CEAM () (Atkinson, T.H.; Equihua M., A. 1985: 238)] [FSCA (4) (Atkinson, T.H.; Equihua M., A. 1985: 238)]
42->	México, Puebla, Puebla	[Unknown () (Wood, S.L. 1982: 839)]
43->	México, Puebla, Texmelucan	[unknown (1) (Wood, S.L. 1982: 839)]
44->	México, Tlaxcala, Calpulalpan; 18/IV/1974; Cibrián T., D.; Pinaceae: Pinus leiophylla	[UACC () (Atkinson, T.H.; Equihua M., A. 1988: 102)]
45->	United States, Arizona, Camp geronimo; 2/VI/1941; Grassman, P.C.	[UAIC (1) (University of Arizona Insect Collection 2016: 0)]
46->	United States, Arizona, Chiricahua Mtns, Idlewild Camp; 25/VII/1980; Turnbow, R.H.; trap sugar bait	[UGCA (1) (TH Atkinson-1988, unpublished)]
47->	United States, Arizona, Chiricahua Mtns.	[unknown (1) (Wood, S.L. 1982: 839)]
48->	United States, Arizona, Flagstaff	[unknown (1) (Wood, S.L. 1982: 839)]
49->	United States, Arizona, Flagstaff; 23/IV/2009; DeGomez, T.; trap alpha pinene + ethanol	[unknown (1) (EDRR 2011: 0)]
50->	United States, Arizona, Jacob's Lake	[unknown (1) (Wood, S.L. 1982: 839)]
51->	United States, Arizona, John Hands Dam, Chiricahua Mtns.; 29/VI/1978; McCleve, S.	[UAIC (1) (University of Arizona Insect Collection 2016: 0)]
52->	United States, Arizona, Kohl's Ranch; 2/VI/1942; Grassman, P.C.	[UAIC (2) (University of Arizona Insect Collection 2016: 0)]
53->	United States, Arizona, Pinal Mtns.	[unknown (1) (Wood, S.L. 1982: 839)]
54->	United States, Arizona, Portal, S.W. Res. Sta., 1.9 mi NW; 28/VIII/1976; Andrews, F.G.; Eichlin, T.; trap: light ultraviolet	[CSCA (1) (TH Atkinson-2014, unpublished)]
55->	United States, Arizona, Prescott N.F.	[unknown (1) (Wood, S.L. 1982: 839)]
56->	United States, Arizona, Williams	[Unknown () (Wood, S.L. 1982: 839)]
57->	United States, Arizona, Williams	[unknown (1) (Wood, S.L. 1982: 839)]
58->	United States, California, Anderson; 18/IV/2007; Seybold, S.J.; Owen, D.; trap alpha pinene + ethanol	[unknown (1) (EDRR 2011: 0)]
59->	United States, California, Big Creek Shaft Road; 11/VI/1972	[CSCA (1) (TH Atkinson-2014, unpublished)]
60->	United States, California, Cleveland Natl. For., Mt. Laguna	[Unknown () (Wood, S.L. 1982: 839)]
61->	United States, California, Crestline; 17/V/2012; Coleman, T.W.; trap: Lindgren Funnel alpha pinene + ethanol	[unknown (1) (EDRR 2014: 0)]
62->	United States, California, El Dorado Co.	[unknown (1) (Bright, D.E., Stark, R.W. 1973: 77)]
63->	United States, California, Forest Falls; 6/V/2011; Coleman, T.; trap alpha pinene + ethanol	[unknown (1) (EDRR 2011: 0)]
64->	United States, California, Fortuna; VI/2001; Spadoni, R.; trap: Lindgren Funnel	[CSCA (1) (TH Atkinson-2014, unpublished)]
65->	United States, California, Groveland; 9/VIII/2012; Bulaon, B.; trap: Lindgren Funnel alpha pinene + ethanol	[unknown (1) (EDRR 2014: 0)]
66->	United States, California, Inyo Co.	[unknown (1) (Bright, D.E., Stark, R.W. 1973: 77)]
67->	United States, California, Jackson Creek	[unknown (1) (Wood, S.L. 1982: 839)]
68->	United States, California, Jenks Lake; 6/V/2011; Coleman, T.; trap alpha pinene + ethanol	[unknown (2) (EDRR 2011: 0)]
69->	United States, California, Lake Co.	[unknown (1) (Bright, D.E., Stark, R.W. 1973: 77)]
70->	United States, California, Lake McK.	[Unknown () (Wood, S.L. 1982: 839)]
71->	United States, California, Los Angeles Co	[unknown (1) (Bright, D.E., Stark, R.W. 1973: 77)]
72->	United States, California, Los Olivos	[unknown (1) (Wood, S.L. 1982: 839)]
73->	United States, California, Madera Co.	[unknown (1) (Wood, S.L. 1982: 839)]
74->	United States, California, Mammoth Lakes; 29/VI/2012; Bulaon, B.; trap: Lindgren Funnel alpha pinene + ethanol	[unknown (1) (EDRR 2014: 0)]
75->	United States, California, Mariposa Co.	[unknown (1) (Bright, D.E., Stark, R.W. 1973: 77)]
76->	United States, California, Mt. Hamilton; 26/V/1972	[CSCA (1) (TH Atkinson-2014, unpublished)]
77->	United States, California, Mt. Laguna	[unknown (1) (Wood, S.L. 1982: 839)]
78->	United States, California, Napa Co.	[unknown (1) (Bright, D.E., Stark, R.W. 1973: 77)]
79->	United States, California, Pinecrest	[unknown (1) (Wood, S.L. 1982: 839)]
80->	United States, California, Redding; 20/VII/2012; Snyder, C.; trap: Lindgren Funnel alpha pinene + ethanol	[unknown (1) (EDRR 2014: 0)]
81->	United States, California, Sagehen Creek; 15/VII/1970; Chandler, D.S.	[UAIC (1) (University of Arizona Insect Collection 2016: 0)]
82->	United States, California, San Bernadino; 12/VII/2013; Coleman, T.W.; trap: Lindgren Funnel alpha pinene + ethanol	[unknown (1) (EDRR 2014: 0)]
83->	United States, California, San Luis Obispo; 19/V/2011; Camilli, K.; trap alpha pinene + ethanol	[unknown (1) (EDRR 2011: 0)]
84->	United States, California, San Luis Obispo; 5/VII/2013; Camilli, K.; trap: Lindgren Funnel ethanol	[unknown (1) (EDRR 2014: 0)]
85->	United States, California, Santa Barbara Co.	[Unknown () (Wood, S.L. 1982: 839)]
86->	United States, California, Santa Barbara Co.	[unknown (1) (Wood, S.L. 1982: 839)]
87->	United States, California, Santa Clara Co.	[unknown (1) (Bright, D.E., Stark, R.W. 1973: 77)]
88->	United States, California, Santa Rosa	[unknown (1) (Wood, S.L. 1982: 839)]
89->	United States, California, Sequoia N.F.	[unknown (1) (Wood, S.L. 1982: 839)]
90->	United States, California, Sequoia National Park; VIII/1929; McClay, A.T.	[UAIC (4) (University of Arizona Insect Collection 2016: 0)]
91->	United States, California, Sequoia Natl. Forest	[Unknown () (Wood, S.L. 1982: 839)]
92->	United States, California, Shasta Natl. Forest	[Unknown () (Wood, S.L. 1982: 839)]
93->	United States, California, Skyforest; 5/V/2008; Coleman, T.; trap alpha pinene + ethanol	[unknown (1) (EDRR 2011: 0)]
94->	United States, California, Susanville; 19/VII/2012; Cluck, D.; trap: Lindgren Funnel alpha pinene + ethanol	[unknown (1) (EDRR 2014: 0)]
95->	United States, California, Susanville; 17/VII/2014; Cluck, D.; trap: Lindgren Funnel exotic ips lure	[unknown (1) (EDRR 2014: 0)]
96->	United States, California, Sylvania	[unknown (1) (Wood, S.L. 1982: 839)]
97->	United States, California, Woodland; 2/VI/2011; Smith, T.; trap alpha pinene + ethanol	[unknown (1) (EDRR 2011: 0)]
98->	United States, Colorado, Allenspark; 14/VI/2011; Daniel, Z.; trap alpha pinene + ethanol	[unknown (1) (EDRR 2011: 0)]
99->	United States, Colorado, Allenspark; 16/VI/2012; Daniel, Z.; trap: Lindgren Funnel alpha pinene + ethanol	[unknown (1) (EDRR 2014: 0)]
100->	United States, Colorado, Blackhawk; 1/VI/2012; Daniel, Z.; trap: Lindgren Funnel exotic ips lure	[unknown (1) (EDRR 2014: 0)]
101->	United States, Colorado, Boulder; 13/VI/2011; Daniel, Z.; trap alpha pinene + ethanol	[unknown (1) (EDRR 2011: 0)]
102->	United States, Colorado, Caribou; 16/VI/2012; Daniel, Z.; trap: Lindgren Funnel ethanol	[unknown (1) (EDRR 2014: 0)]
103->	United States, Colorado, Carter Lake; 3/VI/2012; Rowley, T.; trap: Lindgren Funnel alpha pinene + ethanol	[unknown (1) (EDRR 2014: 0)]
104->	United States, Colorado, Colorado Springs; 25/IV/2007; Aguayo, I.; trap alpha pinene + ethanol	[unknown (1) (EDRR 2011: 0)]
105->	United States, Colorado, Fort Collins; 14/VI/2011; Rowley, T.; trap alpha pinene + ethanol	[unknown (2) (EDRR 2011: 0)]
106->	United States, Colorado, Fort Collins; 2/VII/2012; Rowley, T.; trap: Lindgren Funnel alpha pinene + ethanol	[unknown (1) (EDRR 2014: 0)]
107->	United States, Colorado, Fort Collins; 2/VII/2012; Rowley, T.; trap: Lindgren Funnel ethanol	[unknown (1) (EDRR 2014: 0)]
108->	United States, Colorado, Franktown; 11/IV/2007; Aguayo, I.; trap alpha pinene + ethanol	[unknown (1) (EDRR 2011: 0)]
109->	United States, Colorado, Gilpin; 16/VI/2011; Daniel, Z.; trap alpha pinene + ethanol	[unknown (1) (EDRR 2011: 0)]
110->	United States, Colorado, Livermore; 14/VI/2011; Rowley, T.; trap alpha pinene + ethanol	[unknown (0) (EDRR 2011: 0)]
111->	United States, Colorado, Loveland; 13/VI/2011; Rowley, T.; trap alpha pinene + ethanol	[unknown (1) (EDRR 2011: 0)]
112->	United States, Colorado, Mesa Verde N.P.; 19/VIII/1978	[LSAM (1) (TH Atkinson-1988, unpublished)]
113->	United States, Colorado, Montrose; 1/VI/2007; Hohenstein, J.; trap alpha pinene + ethanol	[unknown (1) (EDRR 2011: 0)]
114->	United States, Colorado, N Cheyenne Canyon	[unknown (1) (Wood, S.L. 1982: 839)]
115->	United States, Colorado, Nederland; 14/VI/2011; Daniel, Z.; trap alpha pinene + ethanol	[unknown (1) (EDRR 2011: 0)]
116->	United States, Colorado, Red Feather Lakes; 14/VI/2011; Rowley, T.; trap alpha pinene + ethanol	[unknown (1) (EDRR 2011: 0)]
117->	United States, Colorado, Stove Praire; 20/VIII/2012; Rowley, T.; trap: Lindgren Funnel exotic ips lure	[unknown (1) (EDRR 2014: 0)]
118->	United States, District of Columbia, Washington	[USNM (1: holotype, female) (Bright, D.E. 1968: 467) (Wood, S.L. 1982: 839) (Hopkins, A.D. 1915: 65) (TH Atkinson-2017, unpublished)]
119->	United States, Florida, Collier; 26/III/2007	[MSUC (1) (Anonymous, 2010: 0)]
120->	United States, Florida, Homestead; 9/VII/2013; Derksen, A.; trap: Lindgren Funnel alpha pinene + ethanol	[unknown (1) (EDRR 2014: 0)]
121->	United States, Georgia, Athens	[Unknown () (Bright, D.E. 1968: 467)]
122->	United States, Georgia, Dunwoody	[Unknown () (Bright, D.E. 1968: 467)]
123->	United States, Georgia, St. George; 7/IV/2011; Price, T.; trap alpha pinene + ethanol	[unknown (1) (EDRR 2011: 0)]
124->	United States, Idaho, 10 km S Howe; Pinaceae: Pseudotsuga menziesii	[Unknown () (Furniss, M.M., Johnson, J.B. 1987: 492)]
125->	United States, Idaho, Moscow Mtn.	[unknown (1) (Wood, S.L. 1982: 839)]
126->	United States, Idaho, Stone Airport, Boise River	[Unknown () (Furniss, M.M., Johnson, J.B. 1987: 492)]
127->	United States, Louisiana, Pineville; 3/IV/2012; Haley, T.; trap: Lindgren Funnel ethanol	[unknown (1) (EDRR 2014: 0)]
128->	United States, Louisiana, Vidalia; 20/III/2012; Haley, T.; trap: Lindgren Funnel ethanol	[unknown (1) (EDRR 2014: 0)]
129->	United States, Maine, Sanford; 21/III/2006; Searles, W.; trap alpha pinene + ethanol	[unknown (1) (EDRR 2011: 0)]
130->	United States, Maryland, Beltsville	[unknown (1) (Wood, S.L. 1982: 839)]
131->	United States, Maryland, Snow Hill	[unknown (1) (Wood, S.L. 1982: 839)]
132->	United States, Mississippi, Adaton, 3 mi W; 9/VI/1997; Schieffer, T.L.; trap: Lindgren Funnel turpentine	[MEM (13) ( : 1)]
133->	United States, Mississippi, Burnsville, 5 mi. S ; Tisdale, R.A.; trap: Lindgren Funnel turpentine + frontalin	[MEM (5) ( : 1)]
134->	United States, Mississippi, M.S.U. Forest T16N, R14E, Sec.5S; 1994; Tisdale, R.A.; trap: Lindgren Funnel turpentine + frontalin	[MEM (5) ( : 1)]
135->	United States, Mississippi, Starkville; 22/VII/1981; Cross, W.H.; trap: light ultraviolet	[MEM (1) ( : 1)]
136->	United States, Montana, Hamilton; 16/IV/2009; Toone, C.; trap alpha pinene + ethanol	[unknown (1) (EDRR 2011: 0)]
137->	United States, Montana, Montana (state)	[unknown (1) (Wood, S.L. 1982: 839)]
138->	United States, Nebraska, Valentine; 8/IV/2008; Harrell, M.; trap alpha pinene + ethanol	[unknown (1) (EDRR 2011: 0)]
139->	United States, Nevada, Las V egas; 21/IV/2009; Knight, J.B.; trap alpha pinene + ethanol	[unknown (1) (EDRR 2011: 0)]
140->	United States, Nevada, Las Vegas; 21/IV/2009; Knight, J.B.; trap alpha pinene + ethanol	[unknown (1) (EDRR 2011: 0)]
141->	United States, Nevada, Reno; 22/IV/2009; Little, R.; trap alpha pinene + ethanol	[unknown (3) (EDRR 2011: 0)]
142->	United States, Nevada, Round Hill; 28/IV/2009; Durham, G.; trap alpha pinene + ethanol	[unknown (3) (EDRR 2011: 0)]
143->	United States, New Jersey, Buena	[Unknown () (Bright, D.E. 1968: 467)]
144->	United States, New Mexico, Cloudcroft	[Unknown () (Wood, S.L. 1982: 839)]
145->	United States, New Mexico, Cloudcroft	[unknown (1) (Wood, S.L. 1982: 839)]
146->	United States, New Mexico, Las Vegas Hot Springs	[Unknown () (Wood, S.L. 1982: 839)]
147->	United States, New Mexico, Lone Mountain Ranch; 2/VIII/2005; Delph, R.; trap: pitfall	[CPMAB (3) (TH Atkinson-2021, unpublished)]
148->	United States, New Mexico, Lone Mountain Ranch; 9/VI/2006; Delph, R.; trap: pitfall	[CPMAB (1) (TH Atkinson-2021, unpublished)]
149->	United States, New Mexico, Los Alamos	[unknown (1) (Wood, S.L. 1982: 839)]
150->	United States, New Mexico, Los Alamos; 7/V/2008; Sandoval, S.; trap alpha pinene + ethanol	[unknown (1) (EDRR 2011: 0)]
151->	United States, New Mexico, Meek	[unknown (1) (Wood, S.L. 1982: 839)]
152->	United States, New Mexico, Santa Fe NF; 5/VIII/2005; Delph, R.; trap: pitfall	[CPMAB (1) (TH Atkinson-2021, unpublished)]
153->	United States, New York, Commack; 4/VI/2013; Denham, J.; Cancelliere, J.; trap: Lindgren Funnel exotic ips lure	[unknown (1) (EDRR 2014: 0)]
154->	United States, New York, Long Island; 25/VII/1898; DuDu Vall, N.R.; Pinaceae: Pinus sp.	[USNM (1: holotype, female) (Bright, D.E. 1968: 467) (Wood, S.L. 1982: 839) (Hopkins, A.D. 1915: 65) (TH Atkinson-2017, unpublished)]
155->	United States, North Carolina, Bent Creek	[unknown (1) (Wood, S.L. 1982: 839)]
156->	United States, North Carolina, Charlotte; 12/IV/2011; Heath; Lawing; trap alpha pinene + ethanol	[unknown (1) (EDRR 2011: 0)]
157->	United States, North Carolina, Dudley; 12/IV/2011; Blaedow, R.; trap alpha pinene + ethanol	[unknown (2) (EDRR 2011: 0)]
158->	United States, North Carolina, McDowell Co.; 1977; Weber, B.C.; trap: window trap	[unknown (1) (Weber, B.C., MacPherson, J.E. 1991: 47)]
159->	United States, North Carolina, Morrisville; 23/V/2013; Oten; Langston; Moan; trap: Lindgren Funnel exotic ips lure	[unknown (1) (EDRR 2014: 0)]
160->	United States, North Carolina, Raleigh; 12/IV/2011; Blaedow, R.; trap alpha pinene + ethanol	[unknown (1) (EDRR 2011: 0)]
161->	United States, North Carolina, Wilmington; 6/VI/2013; Oten; Langston; Moan; trap: Lindgren Funnel alpha pinene + ethanol	[unknown (1) (EDRR 2014: 0)]
162->	United States, Ohio, Ashland; 29/V/2012; Downs; trap: Lindgren Funnel alpha pinene + ethanol	[unknown (1) (EDRR 2014: 0)]
163->	United States, Ohio, Marietta; 24/V/2012; Downs; trap: Lindgren Funnel alpha pinene + ethanol	[unknown (1) (EDRR 2014: 0)]
164->	United States, Oregon, Bend; 25/IV/2007; Hoffman, J.; trap alpha pinene + ethanol	[unknown (1) (EDRR 2011: 0)]
165->	United States, Oregon, Dever	[unknown (1) (Wood, S.L. 1982: 839)]
166->	United States, Oregon, Pine Cr., Blue Mtns.; 6/VII/1980; Haswell, R.H.	[TAMU (1) (TH Atkinson-2012, unpublished)]
167->	United States, Oregon, Woodburn; 7/IV/2011; Adams, T.; trap alpha pinene + ethanol	[unknown (1) (EDRR 2011: 0)]
168->	United States, Pennsylvania, Franklin; 21/V/2013; Coons, S.; trap: Lindgren Funnel alpha pinene + ethanol	[unknown (1) (EDRR 2014: 0)]
169->	United States, Pennsylvania, Liberty; 4/VI/2012; Laubscher, B.; trap: Lindgren Funnel exotic ips lure	[unknown (1) (EDRR 2014: 0)]
170->	United States, Pennsylvania, Mont Alto	[MCZ (4) (TH Atkinson-1988, unpublished)]
171->	United States, Pennsylvania, Mont Alto	[Unknown () (Bright, D.E. 1968: 467) (Wood, S.L. 1982: 839)]
172->	United States, Pennsylvania, Prospect Hill / SGL 61 ; 6/IV/2006; Laubscher, B.; trap alpha pinene + ethanol	[unknown (1) (EDRR 2011: 0)]
173->	United States, Pennsylvania, SGL 247; 6/IV/2006; Williams, J.; trap alpha pinene + ethanol	[unknown (1) (EDRR 2011: 0)]
174->	United States, Pennsylvania, SGL 72; 6/IV/2006; Williams, J.; trap alpha pinene + ethanol	[unknown (1) (EDRR 2011: 0)]
175->	United States, Pennsylvania, SGL 72 ; 6/IV/2006; Williams, J.; trap alpha pinene + ethanol	[unknown (2) (EDRR 2011: 0)]
176->	United States, Pennsylvania, Wildwood Lake Sanctuary ; 29/III/2006; Spichiger, S.; trap alpha pinene + ethanol	[unknown (1) (EDRR 2011: 0)]
177->	United States, Rhode Island, Bristol; 20/IV/2009; Duguay, L.; trap alpha pinene + ethanol	[unknown (2) (EDRR 2011: 0)]
178->	United States, South Carolina, Clemson	[unknown (1) (Wood, S.L. 1982: 839)]
179->	United States, South Dakota, Black Hills; 1/IX/1901; Hopkins, A.D.; Pinaceae: Pinus scopulorum	[USNM (1: holotype, female) (Wood, S.L. 1982: 839) (Hopkins, A.D. 1915: 66) (TH Atkinson-2017, unpublished)]
180->	United States, South Dakota, Pringle	[Unknown () (Wood, S.L. 1982: 839)]
181->	United States, South Dakota, Rapid City; 8/V/2009; Halverson; trap alpha pinene + ethanol	[unknown (1) (EDRR 2011: 0)]
182->	United States, Utah, Fairview; 15/V/2007; Burfitt; trap alpha pinene + ethanol	[unknown (1) (EDRR 2011: 0)]
183->	United States, Utah, Kamas; 6/IV/2007; Burfitt; trap alpha pinene + ethanol	[unknown (1) (EDRR 2011: 0)]
184->	United States, Utah, Logan Canyon; 1946; Wood, S.L.; Pinaceae: Pinus contorta	[USNM (1) (Wood, S.L. 1982: 839) (Wood, S.L. 1951: 128)]
185->	United States, Utah, Provo	[unknown (1) (Wood, S.L. 1982: 839)]
186->	United States, Utah, Provo Canyon	[unknown (1) (Wood, S.L. 1982: 839)]
187->	United States, Utah, Silver Fork; 17/VII/2007; Burfitt; trap alpha pinene + ethanol	[unknown (1) (EDRR 2011: 0)]
188->	United States, Virginia, Falls Church	[unknown (1) (Wood, S.L. 1982: 839)]
189->	United States, West Virginia, Lakin; 27/II/2006; Thaxton, J.; Lowman, A.; trap alpha pinene + ethanol	[unknown (1) (EDRR 2011: 0)]
190->	United States, West Virginia, New Manchester; 1/VI/2006; Braud, R.; trap alpha pinene + ethanol	[unknown (1) (EDRR 2011: 0)]
191->	United States, West Virginia, Romney; 13/III/2006; Drosselmeyer, W.; trap alpha pinene + ethanol	[unknown (1) (EDRR 2011: 0)]