Bark and Ambrosia Beetles of

SYNONYMY

• Debus emarginatus (Eichhoff 1878) Mem. Soc. Roy. Sci. Liege 8(2): 510
• Xyleborus emarginatus Eichhoff 1878. Mem. Soc. Roy. Sci. Liege 8(2): 510
• Xyleborus exesus Blandford 1894. Trans. R. Entomol. Soc. Lond. 1894: 119
• Xyleborus chinchonae Veen 1897. Leyden Mus. Notes 19: 135
• Xyleborus cordatus Hagedorn 1910. Dtsch. Entomol. Z. 1910: 12
• Coptoborus palmeri Hopkins 1915. U.S. Dep. Agric. Tech. Bull. (99): 54
• Coptoborus terminaliae Hopkins 1915. U.S. Dep. Agric. Tech. Bull. (99): 54
• Xyleborus emarginatus semicircularis Schedl 1973. Papua N.G. Agric. J. 24(3): 92

DISTRIBUTION.

Numbers in parentheses after each geographic unit are the number of distinct collection events in the database for that unit. For exotic species generally only countries are listed for localities outside the New World. For further information on published sources of distribution, check the REFERENCES section.

• Asia (27)
  Borneo (1): no state / prov. data (1); China (5): Fujian (1), Shaanxi (1), Sichuan (1), Taiwan (1), Xizang (Tibet) (1); India (1): no state / prov. data (1); Indonesia (5): Java (2), Maluku (1), Sumatra (2); Laos (1): no state / prov. data (1); Malaysia (3): no state / prov. data (1), Sarawak (2); Myanmar (1): no state / prov. data (1); Phillipines (2): Luzon (1), unknown (1); Sri Lanka (1): no state / prov. data (1); Thailand (6): Chiang Mai (1), Nakhon Ratchasima (1), Nakhon Sri Thammarat (1), Songkhla (1), Surat Thani (1), Trang (1); Vietnam (1): no state / prov. data (1);
• Eurasia (1)
  Japan (1): no state / prov. data (1);
• Oceania (3)
  Australia (1): no state / prov. data (1); Papua New Guinea (1): no state / prov. data (1); Solomon Is (1): unknown (1);
• UNK (24)
  (24): no state / prov. data (24);

HOSTS

Numbers in parentheses after each host family, genus, or collection method are the number of distinct collection events in the database for that host or method.

• No host information (31);
• Anacardiaceae (1): Mangifera (1): foetida (1);
• Combretaceae (1): Terminalia (1): sp. (1);
• Dipterocarpaceae (3): Balanocarpus (1): heimii (1); Dipterocarpus (1): baudii (1); Shorea (1): leprosula (1);
• Fagaceae (5): Castanea (1): argentea (1); Castanopsis (2): no species data (1), sp. (1); Pasania (1): cuspidata (1); Quercus (1): sp. (1);
• Gentianaceae (1): Fagraea (1): gigantea (1);
• Leguminosae (3): Albizzia (1): falcata (1); Azfelia (1): palembanica (1); Koompassia (1): excelsa (1);
• Malvaceae (1): Durio (1): zibethinus (1);
• Moraceae (1): Ficus (1): sp. (1);
• Pinaceae (3): Abies (1): fabri (1); Pinus (2): tabulaeformis (1), yunnanensis (1);
• Rubiaceae (2): Cinchona (1): sp. (1); Sarcocephalus (1): cordatus (1);
• Salicaceae (1): Populus (1): sp. (1);
• Sapotaceae (1): Palaquium (1): sp. (1);
• Symplocaceae (1) Symplocos (1): sp. (1);

REFERENCES

The following are important recent monographs, catalogs, and supplements to catalogs that refer to this species. The specific page on which the reference is made is shown in pink at the end of the reference. In the case of Wood (1982) and Wood & Bright (1992) clicking on the reference page will link to a digital version of the work in question.

• Wood, S.L., Bright,D.E. 1992. A catalog of Scolytidae and Platypodidae (Coleoptera), Part 2. Taxonomic Index. Great Basin Nat. Mem. 13:1-1553 (vol. A, B). [733]

The following are references from which host and distribution data have been input into the database. If one of the above monographs or catalogs also appears in this list, it means that most relevant collection event data have been included.

1. Beaver, R.A.; Sittichaya, W.; Liu, L.Y. 2014. A Synopsis of the Scolytine Ambrosia Beetles of Thailand (Coleoptera: Curculionidae: Scolytinae). Zootaxa 3875(1): 1-82.
2. Bright, D.E. 2000. Scolytidae (Coleoptera) of Gunung Mulu National Park, Sarawak, Malaysia, with ecological notes and descriptions of six new species. Serangga 5: 41-85.
3. Hopkins, A.D. 1915. Classification of the Cryphalinae with descriptions of new genera and species. USDA Ent. Comm. Report (99): 1-75.
4. Wood, S.L.; Bright, D.E. 1992. A catalog of Scolytidae and Platypodidae (Coleoptera), Part 2. Taxonomic Index (Volumes A,B). Great Basin Nat. Mem. 13: 1-1553.

Maps automatically open at the center of the plotted points and the scale is set to encompass all map points. Maps can be resized (scale bar at upper left) and the center moved (place cursor over map and drag) to see other parts of the distribution of the species. Clicking on a map marker will pull up collection event data and a literature citation if present. If any errors are found, please refer to the "series code" which is a unique identifier for a database record in any communications). Coordinates have not been entered for all collection records. Localities outside the New World are not plotted, even though they are listed in the distribution summary and in the table of records.

No.	Collection data	Publication
Asia
1->	Borneo, no state / prov. data, 'Borneo' [Wood & Bright 1992 confounded Malaysian and Indone	[unknown (1) (Wood, S.L.; Bright, D.E. 1992: 733)]
2->	China, Fujian, Fujian	[unknown (1) (Wood, S.L.; Bright, D.E. 1992: 733)]
3->	China, Shaanxi , Shanxi	[unknown (1) (Wood, S.L.; Bright, D.E. 1992: 733)]
4->	China, Sichuan, Sichuan	[unknown (1) (Wood, S.L.; Bright, D.E. 1992: 733)]
5->	China, Taiwan, Taiwan	[unknown (1) (Beaver, R.A.; Sittichaya, W.; Liu, L.Y. 2014: 42)]
6->	China, Xizang (Tibet), Xizang	[unknown (1) (Wood, S.L.; Bright, D.E. 1992: 733)]
7->	India, no state / prov. data, Nicobar Islands	[unknown (1) (Beaver, R.A.; Sittichaya, W.; Liu, L.Y. 2014: 42)]
8->	Indonesia, Java, Java	[RMNH (1: syntype(s), female) (Wood, S.L.; Bright, D.E. 1992: 733)]
9->	Indonesia, Java, Mount Salak; 3/V/1909; Bryant; Palmer	[USNM (1: holotype, female) (Wood, S.L.; Bright, D.E. 1992: 664) (TH Atkinson-2018, unpublished) (Hopkins, A.D. 1915: 54)]
10->	Indonesia, Maluku, Moluccas	[unknown (1) (Wood, S.L.; Bright, D.E. 1992: 733)]
11->	Indonesia, Sumatra, Mentawaii	[ZMHB (1: syntype(s), female) (Wood, S.L.; Bright, D.E. 1992: 733)]
12->	Indonesia, Sumatra, Sumatra	[unknown (1) (Wood, S.L.; Bright, D.E. 1992: 733)]
13->	Laos, no state / prov. data, Laos	[unknown (1) (Wood, S.L.; Bright, D.E. 1992: 733)]
14->	Malaysia, no state / prov. data, Malaysia	[unknown (1) (Wood, S.L.; Bright, D.E. 1992: 733)]
15->	Malaysia, Sarawak, Gunung Mulu NP; 1978; Hanski, I.; Hammond, P.	[BMNH (1) (Bright, D.E. 2000: 69)]
16->	Malaysia, Sarawak, Gunung Mulu NP; 1978; Hanski, I.; Hammond, P.	[BMNH (1) (Bright, D.E. 2000: 69)]
17->	Myanmar, no state / prov. data, Burma	[ZUMH (0: holotype, female) (Wood, S.L.; Bright, D.E. 1992: 733)]
18->	Phillipines, Luzon, Pagbilao; 4/VIII/1903; Webb, J.L.; Combretaceae: Terminalia sp.	[USNM (1: holotype, female) (Wood, S.L.; Bright, D.E. 1992: 664) (TH Atkinson-2018, unpublished) (Hopkins, A.D. 1915: 54)]
19->	Phillipines, unknown, Phillipines (country)	[unknown (1) (Wood, S.L.; Bright, D.E. 1992: 733)]
20->	Sri Lanka, no state / prov. data, Sri Lanka	[unknown (1) (Wood, S.L.; Bright, D.E. 1992: 733)]
21->	Thailand, Chiang Mai, Chiang Mai (province)	[unknown (1) (Beaver, R.A.; Sittichaya, W.; Liu, L.Y. 2014: 42)]
22->	Thailand, Nakhon Ratchasima, Nakhon Ratchasima (province)	[unknown (1) (Beaver, R.A.; Sittichaya, W.; Liu, L.Y. 2014: 42)]
23->	Thailand, Nakhon Sri Thammarat, Nakhon Sri Thammarat (province)	[unknown (1) (Beaver, R.A.; Sittichaya, W.; Liu, L.Y. 2014: 42)]
24->	Thailand, Songkhla, Songkhla (province)	[unknown (1) (Beaver, R.A.; Sittichaya, W.; Liu, L.Y. 2014: 42)]
25->	Thailand, Surat Thani, Surat Thani (province)	[unknown (1) (Beaver, R.A.; Sittichaya, W.; Liu, L.Y. 2014: 42)]
26->	Thailand, Trang, Trang (province)	[unknown (1) (Beaver, R.A.; Sittichaya, W.; Liu, L.Y. 2014: 42)]
27->	Vietnam, no state / prov. data, Vietnam	[unknown (1) (Wood, S.L.; Bright, D.E. 1992: 733)]
Eurasia
28->	Japan, no state / prov. data, Miyanoshita	[BMNH (1: holotype, female) (Wood, S.L.; Bright, D.E. 1992: 734)]
Oceania
29->	Australia, no state / prov. data, Australia	[unknown (1) (Wood, S.L.; Bright, D.E. 1992: 733)]
30->	Papua New Guinea, no state / prov. data, Porotop Mission Station, Western Highlands	[CSIRO (1: holotype, female) (Wood, S.L.; Bright, D.E. 1992: 733)]
31->	Solomon Is, unknown, Solomon Is (country)	[unknown (1) (Beaver, R.A.; Sittichaya, W.; Liu, L.Y. 2014: 42)]
UNK
32->	, no state / prov. data	[TAMU () (TH Atkinson-, unpublished)]
33->	, no state / prov. data; Pinaceae: Abies fabri	[unknown (0) (Wood, S.L.; Bright, D.E. 1992: 733)]
34->	, no state / prov. data; Leguminosae: Albizzia falcata	[unknown (0) (Wood, S.L.; Bright, D.E. 1992: 733)]
35->	, no state / prov. data; Dipterocarpaceae: Balanocarpus heimii	[unknown (0) (Wood, S.L.; Bright, D.E. 1992: 733)]
36->	, no state / prov. data; Fagaceae: Castanea argentea	[unknown (0) (Wood, S.L.; Bright, D.E. 1992: 733)]
37->	, no state / prov. data; Fagaceae: Castanopsis sp.	[unknown (0) (Wood, S.L.; Bright, D.E. 1992: 733)]
38->	, no state / prov. data; Fagaceae: Castanopsis	[unknown (0) (Wood, S.L.; Bright, D.E. 1992: 733)]
39->	, no state / prov. data; Rubiaceae: Cinchona sp.	[unknown (0) (Wood, S.L.; Bright, D.E. 1992: 733)]
40->	, no state / prov. data; Dipterocarpaceae: Dipterocarpus baudii	[unknown (0) (Wood, S.L.; Bright, D.E. 1992: 733)]
41->	, no state / prov. data; Malvaceae: Durio zibethinus	[unknown (0) (Wood, S.L.; Bright, D.E. 1992: 733)]
42->	, no state / prov. data; Gentianaceae: Fagraea gigantea	[unknown (0) (Wood, S.L.; Bright, D.E. 1992: 733)]
43->	, no state / prov. data; Moraceae: Ficus sp.	[unknown (0) (Wood, S.L.; Bright, D.E. 1992: 733)]
44->	, no state / prov. data; Leguminosae: Azfelia palembanica	[unknown (0) (Wood, S.L.; Bright, D.E. 1992: 733)]
45->	, no state / prov. data; Leguminosae: Koompassia excelsa	[unknown (0) (Wood, S.L.; Bright, D.E. 1992: 733)]
46->	, no state / prov. data; Anacardiaceae: Mangifera foetida	[unknown (0) (Wood, S.L.; Bright, D.E. 1992: 733)]
47->	, no state / prov. data; Sapotaceae: Palaquium sp.	[unknown (0) (Wood, S.L.; Bright, D.E. 1992: 733)]
48->	, no state / prov. data; Pinaceae: Pinus tabulaeformis	[unknown (0) (Wood, S.L.; Bright, D.E. 1992: 733)]
49->	, no state / prov. data; Pinaceae: Pinus yunnanensis	[unknown (0) (Wood, S.L.; Bright, D.E. 1992: 733)]
50->	, no state / prov. data; Salicaceae: Populus sp.	[unknown (0) (Wood, S.L.; Bright, D.E. 1992: 733)]
51->	, no state / prov. data; Fagaceae: Quercus sp.	[unknown (0) (Wood, S.L.; Bright, D.E. 1992: 733)]
52->	, no state / prov. data; Rubiaceae: Sarcocephalus cordatus	[unknown (0) (Wood, S.L.; Bright, D.E. 1992: 733)]
53->	, no state / prov. data; Dipterocarpaceae: Shorea leprosula	[unknown (0) (Wood, S.L.; Bright, D.E. 1992: 733)]
54->	, no state / prov. data; Symplocaceae: Symplocos sp.	[unknown (0) (Wood, S.L.; Bright, D.E. 1992: 733)]
55->	, no state / prov. data; Fagaceae: Pasania cuspidata	[unknown (0) (Wood, S.L.; Bright, D.E. 1992: 734)]