Bark and Ambrosia Beetles of

SYNONYMY

• Pityoborus secundus Blackman 1928. NY State Coll. For. Tech. Pub. 25: 146
• Pityoborus tertius Blackman 1942. Proc. USNM 93: 202
• Pityoborus intonsus Wood 1958. Great Basin Nat. 18: 54
• Pityoborus immitus Bright 1972. Can. Ent. 104: 1674
• Pityoborus ramosus Bright 1972. Can. Ent. 104: 1677

DISTRIBUTION.

Numbers in parentheses after each geographic unit are the number of distinct collection events in the database for that unit. For exotic species generally only countries are listed for localities outside the New World. For further information on published sources of distribution, check the REFERENCES section.

Distribution Comments.  Wood (1982) and Wood & Bright (1992) list Texcoco from the state of Veracruz. They also list Chalco from the Fderal District (DF). Both are in the state of Mexico.

• North America (29)
  (4): no state / prov. data (4); México (18): Chiapas (1), Chihuahua (2), Durango (4), Hidalgo (1), México (2), Michoacán (2), Morelos (3), Oaxaca (1), Puebla (1), Tlaxcala (1); United States (7): Arizona (5), New Mexico (1), Utah (1);

HOSTS

Numbers in parentheses after each host family, genus, or collection method are the number of distinct collection events in the database for that host or method.

Host Comments.  In shaded out branches of living trees.

• No host information (15);
• Pinaceae (11): Pinus (11): ayacahuite (1), cooperi (1), engelmanni (1), engelmannii (1), lawsoni (1), leiophylla (1), oocarpa (1), ponderosa (1), pseudostrobus (1), sp. (2);
• trap (3) bottle (1): no details (1); light (1): ultraviolet (1); Lindgren Funnel (1): MPB pheromone (1);

REFERENCES

The following are important recent monographs, catalogs, and supplements to catalogs that refer to this species. The specific page on which the reference is made is shown in pink at the end of the reference. In the case of Wood (1982) and Wood & Bright (1992) clicking on the reference page will link to a digital version of the work in question.

• Wood, S.L. 1982. The bark and ambrosia beetles of North and Central America (Coleoptera: Scolytidae), a taxonomic monograph. Great Basin Nat. Mem. 6:1-1356. [1148]. (data capture complete)
• Wood, S.L., Bright,D.E. 1992. A catalog of Scolytidae and Platypodidae (Coleoptera), Part 2. Taxonomic Index. Great Basin Nat. Mem. 13:1-1553 (vol. A, B). [968]

The following are references from which host and distribution data have been input into the database. If one of the above monographs or catalogs also appears in this list, it means that most relevant collection event data have been included.

1. . . .
2. Atkinson, T.H.; Saucedo C.; E, Martínez F.; E, Burgos S., A 1986. Coleópteros Scolytidae y Platypodidae asociados con las comunidades vegetales de clima templado y frío en el estado de Morelos. Acta Zool. Mex. 17: 1-58.
3. Lázaro Dzul, M.O.; Equihua Martinez, A.; Romero Nápoles, J.; Gonzalez H., H.; Alvarado, D.; Macías Samaño, J.; Castañeda Vildóla, A.; Atkinson, T,H, 2020. Diversity of Scolytinae (Coleoptera: Curculionidae) at three Sites in the Central Avocado-Producing Region of Michoacán, Mexico. Coleopterists Bull. 74 (2): 454-462.
4. University of Arizona Insect Collection 2016. SCAN: Symbiota Collections of Arthropods Network. http://symbiota4.acis.ufl.edu/scan/portal/collections/misc/collprofiles.php?collid=11.
5. Wood, S.L. 1982. The bark and ambrosia beetles of North and Central America (Coleoptera: Scolytidae), a taxonomic monograph. Great Basin Nat. Mem. 6: 1-1356.

Maps automatically open at the center of the plotted points and the scale is set to encompass all map points. Maps can be resized (scale bar at upper left) and the center moved (place cursor over map and drag) to see other parts of the distribution of the species. Clicking on a map marker will pull up collection event data and a literature citation if present. If any errors are found, please refer to the "series code" which is a unique identifier for a database record in any communications). Coordinates have not been entered for all collection records. Localities outside the New World are not plotted, even though they are listed in the distribution summary and in the table of records.

No.	Collection data	Publication
North America
1->	, no state / prov. data; Pinaceae: Pinus ayacahuite	[unknown (0) (Wood, S.L. 1982: 1148)]
2->	, no state / prov. data; Pinaceae: Pinus leiophylla	[unknown (0) (Wood, S.L. 1982: 1148)]
3->	, no state / prov. data; Pinaceae: Pinus ponderosa	[unknown (0) (Wood, S.L. 1982: 1148)]
4->	, no state / prov. data; Pinaceae: Pinus pseudostrobus	[unknown (0) (Wood, S.L. 1982: 1148)]
5->	México, Chiapas, Angel Corzo; 4/XI/1984; Bermúdez, N.; Pinaceae: Pinus oocarpa	[CEAM () (Atkinson, T.H.; Equihua M., A. 1988: 94)]
6->	México, Chihuahua, Goachachic; 29/IV/1981; Furniss, M.M.; Cibrían T., D.; Pinaceae: Pinus cooperi	[UACC () (Atkinson, T.H.; Equihua M., A. 1988: 94)]
7->	México, Chihuahua, Las Varas, 7 mi W; 24/IV/1981; Furniss, M.M.; Cibrían T., D.; Pinaceae: Pinus engelmannii	[UACC () (Atkinson, T.H.; Equihua M., A. 1988: 94)]
8->	México, Durango, Durango, 28 mi SW; 19/VIII/1970; Johnson, C.D.	[CPMAB (1) (TH Atkinson-2021, unpublished)]
9->	México, Durango, El Salto, 17 km W	[unknown (1) (Wood, S.L. 1982: 1148)]
10->	México, Durango, El Salto, 37 mi (68 km) SW; 18/VI/1971; Bright, D.E.; Pinaceae: Pinus engelmanni	[CNC (1: holotype, female) (Wood, S.L. 1982: 1148)]
11->	México, Durango, El Salto, 5 km W	[unknown (1) (Wood, S.L. 1982: 1148)]
12->	México, Hidalgo, Jacala, 17 km N	[unknown (1) (Wood, S.L. 1982: 1148)]
13->	México, México, Chalco	[USNM (1: holotype, female) (Wood, S.L. 1982: 1148) (TH Atkinson-2017, unpublished)]
14->	México, México, Texcoco	[unknown (1) (Wood, S.L. 1982: 1148)]
15->	México, Michoacán, Nuevo San Juan Parangarícutiro ; VII/2016; Lázaro Dzul, M.O.; trap: bottle	[unknown (1) (Lázaro Dzul, M.O.; Equihua Martinez, A.; Romero Nápoles, J.; Gonzalez H., H.; Alvarado, D.; Macías Samaño, J.; Castañeda Vildóla, A.; Atkinson, T,H, 2020: 0)]
16->	México, Michoacán, Volcan Paricutin	[unknown (1) (Wood, S.L. 1982: 1148)]
17->	México, Morelos, Chamilpa; trap: light ultraviolet	[UAEM () (Atkinson, T.H.; Saucedo C.; E, Martínez F.; E, Burgos S., A 1986: 36)]
18->	México, Morelos, El Capulin; Burgos S., A.; Pinaceae: Pinus sp.	[CEAM () (Atkinson, T.H.; Saucedo C.; E, Martínez F.; E, Burgos S., A 1986: 36)] [TAMU (2) (Atkinson, T.H.; Saucedo C.; E, Martínez F.; E, Burgos S., A 1986: 36)]
19->	México, Morelos, Tres Marias; Burgos S., A.; Pinaceae: Pinus sp.	[UAEM () (Atkinson, T.H.; Saucedo C.; E, Martínez F.; E, Burgos S., A 1986: 36)]
20->	México, Oaxaca, Suchixtepec, 3.5 mi S, hwy 175; 2/VI/1971; Bright, D.E.; Pinaceae: Pinus lawsoni	[CNC (1: holotype, female) (Wood, S.L. 1982: 1148)]
21->	México, Puebla, Texmelucán, 14 mi (23 km) W	[SEMC (1: holotype, female) (Wood, S.L. 1982: 1148)]
22->	México, Tlaxcala, Temetzontla; 17/VI/1986; Campos B., R.; Cibrián T., D.; trap: Lindgren Funnel MPB pheromone	[TAMU (2) (TH Atkinson-1989, unpublished)]
23->	United States, Arizona, Bear Canyon, Hitchcock Hwy, mi 11; 9/VII/1959; Werner, F.G.; Radford, K.W.; Samuelson, G.A.	[UAIC (4) (University of Arizona Insect Collection 2016: 0)]
24->	United States, Arizona, Lakeside	[unknown (1) (Wood, S.L. 1982: 1148)]
25->	United States, Arizona, Oak Creek Canyon	[unknown (1) (Wood, S.L. 1982: 1148)]
26->	United States, Arizona, Pinetop; 25/VIII/1996; Greeney, H.	[UAIC (1) (University of Arizona Insect Collection 2016: 0)]
27->	United States, Arizona, Prescott N.F.	[unknown (1) (Wood, S.L. 1982: 1148)]
28->	United States, New Mexico, Cloudcroft	[unknown (1) (Wood, S.L. 1982: 1148)]
29->	United States, Utah, La Sal Mtns.	[USNM (1: holotype, female) (Wood, S.L. 1982: 1148) (TH Atkinson-2017, unpublished)]