Bark and Ambrosia Beetles of

SYNONYMY

• Phloeotribus frontalis (Olivier 1795) Entomologie Coleopt. 4(78): 13
• Bostrichus frontalis (Olivier 1795) Entomologie Coleopt. 4(78): 13
• Phthorophloeus frontalis (Olivier 1795) Entomologie Coleopt. 4(78): 13
• Scolytus frontalis Olivier 1795. Entomologie Coleopt. 4(78): 13
• Phloeotribus granicollis (Eichhoff 1868) Berliner Ent. Zeit. 12: 149
• Phloeophthorus granicollis Eichhoff 1868. Berliner Ent. Zeit. 12: 149
• Phloeophthorus moriperda Hopkins 1905. Proc. Ent. Soc. Wash. 7: 77

DISTRIBUTION.

Numbers in parentheses after each geographic unit are the number of distinct collection events in the database for that unit. For exotic species generally only countries are listed for localities outside the New World. For further information on published sources of distribution, check the REFERENCES section.

• North America (163)
  México (3): Guanajuato (1), México (1), Oaxaca (1); United States (160): no state / prov. data (2), Alabama (4), Arkansas (1), Delaware (1), District of Columbia (1), Florida (5), Georgia (14), Illinois (2), Indiana (9), Iowa (3), Kansas (2), Kentucky (2), Louisiana (8), Maryland (4), Michigan (2), Mississippi (4), Missouri (15), Nebraska (4), New Jersey (4), New York (4), North Carolina (9), Ohio (2), Oklahoma (18), Pennsylvania (8), South Carolina (5), Tennessee (4), Texas (10), Virginia (3), West Virginia (10);

HOSTS

Numbers in parentheses after each host family, genus, or collection method are the number of distinct collection events in the database for that host or method.

• No host information (93);
• Cannabaceae (2): Celtis (2): laevigata (1), occidentalis (1);
• Leguminosae (1): Robinia (1): sp. (1);
• Moraceae (15): Ficus (1): carica (1); Maclura (1): pomifera (1); Morus (13): alba (2), rubra (6), sp. (5);
• trap (51): (26): alpha pinene + ethanol (25), sugar bait (1); alpha-scents 4 vane (3): ethanol (3); bottle (1): ethanol (1); canopy (1): no details (1); flight intercept (3): no details (3); light (4): ultraviolet (4); Lindgren Funnel (11): alpha pinene + ethanol (3), ethanol (2), exotic ips lure (6); window trap (2): no details (2);
• Ulmaceae (1) Ulmus (1): sp. (1);

REFERENCES

The following are important recent monographs, catalogs, and supplements to catalogs that refer to this species. The specific page on which the reference is made is shown in pink at the end of the reference. In the case of Wood (1982) and Wood & Bright (1992) clicking on the reference page will link to a digital version of the work in question.

• Wood, S.L. 1982. The bark and ambrosia beetles of North and Central America (Coleoptera: Scolytidae), a taxonomic monograph. Great Basin Nat. Mem. 6:1-1356. [270]. (data capture complete)
• Wood, S.L., Bright,D.E. 1992. A catalog of Scolytidae and Platypodidae (Coleoptera), Part 2. Taxonomic Index. Great Basin Nat. Mem. 13:1-1553 (vol. A, B). [221]

The following are references from which host and distribution data have been input into the database. If one of the above monographs or catalogs also appears in this list, it means that most relevant collection event data have been included.

1. . . .
2. Atkinson, T.H.; Rabaglia,R.J.; Peck,S.J.; Foltz,J.L. 1991. New records of Scolytidae and Platypodidae from the United States and Bahamas. Coleopterists Bull. 45(2): 152-164.
3. Beal, J.A., Massey, C.L. 1945. Bark beetles and ambrosia beetles (Coleoptera: Scolytoidea) with special reference to the species ocurring in North Carolina. Duke Univ. School of Forestry Bull. 10: 1-178.
4. Deyrup, M.A. 1981. Annotated list of Indiana Scolytidae (Coleoptera). Great Lakes Entomol. 14: 1-9.
5. Deyrup, M.A.; Atkinson, T.H. 1987. New records of Scolytidae (Coleoptera) from Indiana and Florida. Great Lakes Entomol. 20: 67-68.
6. EDRR 2011. USFS Early Detection Rapid Response Database. http://www.fs.fed.us/invasivespecies/earlydetection.shtml/.
7. EDRR 2014. USFS Early Detection Rapid Response Database. http://www.fs.fed.us/invasivespecies/earlydetection.shtml/.
8. Hopkins, A.D. 1905. Notes on some Mexican Scolytidae with descriptions of new species. Proc. Ent. Soc. Wash. 7: 71-81.
9. Viloria, Z.; Villanueva, R.T.; Bessin, R.; O'Neal, P.; Ranger, C.M.; Dunwell, W. 2021. Scolytinae in Nursery and Fruit Crops of Western Kentucky and Seasonal Population Patterns of Four Invasive Ambrosia beetles. J. Entomol. Sci. 56 (3): 374-386.
10. Weber, B.C., MacPherson, J.E. 1991. Seasonal Flight Patterns Of Scolytidae (Coleoptera) In Black-Walnut Plantations In North-Carolina And Illinois. Coleopterists Bull. 45(1): 45-56.
11. Wood, S.L. 1982. The bark and ambrosia beetles of North and Central America (Coleoptera: Scolytidae), a taxonomic monograph. Great Basin Nat. Mem. 6: 1-1356.

Maps automatically open at the center of the plotted points and the scale is set to encompass all map points. Maps can be resized (scale bar at upper left) and the center moved (place cursor over map and drag) to see other parts of the distribution of the species. Clicking on a map marker will pull up collection event data and a literature citation if present. If any errors are found, please refer to the "series code" which is a unique identifier for a database record in any communications). Coordinates have not been entered for all collection records. Localities outside the New World are not plotted, even though they are listed in the distribution summary and in the table of records.

No.	Collection data	Publication
North America
1->	México, Guanajuato, Irapuato; Chambon, H.; Moraceae: Morus alba	[USNM (1: holotype) (Wood, S.L. 1982: 270) (Hopkins, A.D. 1905: 77)]
2->	México, México, Chiconcuac; 14/X/1980; Atkinson, T.H.; Moraceae: Morus alba	[CEAM (19) (Atkinson, T.H.; Equihua M., A. 1985: 76) (TH Atkinson-2015, unpublished)]
3->	México, Oaxaca, Hwy 175, 6.3 km N jct hwy 190; 20/VI/2009; Atkinson, T.H.; Moraceae: Morus sp.	[UTIC (4) (TH Atkinson-2009, unpublished)] [TAMU (20) (TH Atkinson-2009, unpublished)] [FSCA (8) (TH Atkinson-2009, unpublished)] [CAS (8) (TH Atkinson-2009, unpublished)] [CEAM (8) (TH Atkinson-2009, unpublished)] [MSUC (8) (TH Atkinson-2009, unpublished)]
4->	United States, no state / prov. data, America Boreale	[USNM (1: lectotype, male) (Wood, S.L. 1982: 270) (TH Atkinson-2018, unpublished)]
5->	United States, no state / prov. data, America Septentrionale	[MNHN (1: holotype) (Wood, S.L. 1982: 270)]
6->	United States, Alabama, Alabama (state)	[unknown () (Wood, S.L. 1982: 270)]
7->	United States, Alabama, Birmingham	[USNM () (TH Atkinson-1988, unpublished)]
8->	United States, Alabama, DC	[unknown () (Wood, S.L. 1982: 270)]
9->	United States, Alabama, Toney; Moraceae: Morus sp.	[USNM () (TH Atkinson-1988, unpublished)]
10->	United States, Arkansas, Little Rock; 4/V/1998; Baldwin; trap: flight intercept	[TAMU (1) (TH Atkinson-2003, unpublished)]
11->	United States, Delaware, Delaware (state)	[MCZ () (TH Atkinson-1988, unpublished)]
12->	United States, District of Columbia, Washington	[USNM () (TH Atkinson-1988, unpublished)]
13->	United States, Florida, Branford; I/1988; Atkinson, T.H.; Moraceae: Morus rubra	[TAMU (20) (TH Atkinson-1989, unpublished)]
14->	United States, Florida, Gainesville; 31/VII/1987; Atkinson, T.H.; Moraceae: Morus rubra	[TAMU (4) (Deyrup, M.A.; Atkinson, T.H. 1987: 68)]
15->	United States, Florida, Gainesville; 22/IV/1977; Atkinson, T.H.; Moraceae: Morus rubra	[TAMU (53) (Deyrup, M.A.; Atkinson, T.H. 1987: 68)] [CEAM (6) (TH Atkinson-2015, unpublished)]
16->	United States, Florida, San Felasco Hammock St Pres; 10/III/1992; Turnbow, R.H.	[RHTC (1) (TH Atkinson-2008, unpublished)]
17->	United States, Florida, San Felasco Hammock St Pres; 15/III/1992; Turnbow, R.H.	[RHTC (18) (TH Atkinson-2008, unpublished)]
18->	United States, Georgia, Athens; 15/III/1982; Monkman, K.; Moraceae: Ficus carica	[UGCA (33) (TH Atkinson-1988, unpublished)]
19->	United States, Georgia, Athens; 17/IV/2014; Barnes, B.; trap: Lindgren Funnel exotic ips lure	[unknown (1) (EDRR 2014: 0)]
20->	United States, Georgia, Athens, 3 mi N; 12/IV/1974; Turnbow, R.H.	[UGCA (1) (TH Atkinson-1988, unpublished)]
21->	United States, Georgia, Athens, 3 mi N; 5/IV/1978; Turnbow, R.H.; trap: light ultraviolet	[UGCA (1) (TH Atkinson-1988, unpublished)]
22->	United States, Georgia, Bishop; 1/IV/2011; Brownell, K.; trap alpha pinene + ethanol	[unknown (1) (EDRR 2011: 0)]
23->	United States, Georgia, Georgia (state)	[unknown () (Wood, S.L. 1982: 270)]
24->	United States, Georgia, Grady Co.; 31/III/; Hudson, W.	[MEFEIS (1) (TH Atkinson-1997, unpublished)]
25->	United States, Georgia, Lithonia; 8/IV/2011; Price, T.; trap alpha pinene + ethanol	[unknown (1) (EDRR 2011: 0)]
26->	United States, Georgia, Milner	[USNM () (TH Atkinson-1988, unpublished)]
27->	United States, Georgia, Watkinsville; 27/IV/2012; Adair, J.; trap: Lindgren Funnel exotic ips lure	[unknown (1) (EDRR 2014: 0)]
28->	United States, Georgia, Whitehall Forest; 13/VII/1979; Turnbow, R.H.; trap sugar bait	[UGCA (6) (TH Atkinson-1988, unpublished)]
29->	United States, Georgia, Winterville; 1/IV/2011; Brownell, K.; trap alpha pinene + ethanol	[unknown (1) (EDRR 2011: 0)]
30->	United States, Georgia, Winterville; 16/IV/2014; Barnes, B.; trap: Lindgren Funnel exotic ips lure	[unknown (1) (EDRR 2014: 0)]
31->	United States, Georgia, Winterville; 18/IV/2013; Barnes, B.; trap: Lindgren Funnel exotic ips lure	[unknown (1) (EDRR 2014: 0)]
32->	United States, Illinois, Alexander Co.; 1989; Weber, B.C.; trap: window trap	[unknown (1) (Weber, B.C., MacPherson, J.E. 1991: 55)]
33->	United States, Illinois, Illinois (state)	[unknown () (Wood, S.L. 1982: 270)]
34->	United States, Indiana; Moraceae: Morus rubra	[unknown (1) (Deyrup, M.A. 1981: 4)]
35->	United States, Indiana, Bloomington; 13/IV/1985; Young, F.N.; trap: light ultraviolet	[FSCA (1) (TH Atkinson-2007, unpublished)]
36->	United States, Indiana, Bloomington; 1/V/1993; Young, F.N.; trap: light ultraviolet	[FSCA (1) (TH Atkinson-2007, unpublished)]
37->	United States, Indiana, Brown Co.	[Unknown () (Deyrup, M.A. 1981: 4)]
38->	United States, Indiana, Clay Co.	[Unknown () (Deyrup, M.A. 1981: 4)]
39->	United States, Indiana, Evansville	[USNM () (TH Atkinson-1988, unpublished)]
40->	United States, Indiana, Indiana (state)	[unknown () (Wood, S.L. 1982: 270)]
41->	United States, Indiana, Parke Co	[Unknown () (Deyrup, M.A. 1981: 4)]
42->	United States, Indiana, Tippecanoe Co.	[Unknown () (Deyrup, M.A. 1981: 4)]
43->	United States, Iowa, Burlington	[MCZ () (TH Atkinson-1988, unpublished)]
44->	United States, Iowa, Iowa (state)	[unknown () (Wood, S.L. 1982: 270)]
45->	United States, Iowa, Iowa City	[USNM () (TH Atkinson-1988, unpublished)]
46->	United States, Kansas, Crestline, 3 mi E, 1 mi N; 21/IV/2009; Salsbury, G.; trap: canopy	[SEMC (2) (TH Atkinson-2020, unpublished)]
47->	United States, Kansas, Kansas (state)	[unknown () (Wood, S.L. 1982: 270)]
48->	United States, Kentucky, Barbourville; 1/V/1965; White, R.E.	[FSCA (15) (TH Atkinson-2014, unpublished)]
49->	United States, Kentucky, Princeton; 2016; trap: bottle ethanol	[unknown (1) (Viloria, Z.; Villanueva, R.T.; Bessin, R.; O'Neal, P.; Ranger, C.M.; Dunwell, W. 2021: 378)]
50->	United States, Louisiana, Alexandria	[USNM () (TH Atkinson-1988, unpublished)]
51->	United States, Louisiana, Alexandria; 1/III/2010; Haley, T.; trap alpha pinene + ethanol	[unknown (3) (EDRR 2011: 0)]
52->	United States, Louisiana, Bayou Sara	[USNM () (TH Atkinson-1988, unpublished)]
53->	United States, Louisiana, Bodcau Wildlife Mmt. Area; 30/V/1966; Riley, E.G.; trap: light ultraviolet	[TAMU (1) (TH Atkinson-2003, unpublished)]
54->	United States, Louisiana, Forest Hill; 1/III/2010; Haley, T.; trap alpha pinene + ethanol	[unknown (1) (EDRR 2011: 0)]
55->	United States, Louisiana, Louisiana (state)	[unknown () (Wood, S.L. 1982: 270)]
56->	United States, Louisiana, Louisiana (state)	[unknown (1) (Wood, S.L. 1982: 271)]
57->	United States, Louisiana, Shreveport; 1/III/2010; Haley, T.; trap alpha pinene + ethanol	[unknown (0) (EDRR 2011: 0)]
58->	United States, Maryland, Frederick; 28/III/2011; Lupp, T.; Firme, C.A.; trap alpha pinene + ethanol	[unknown (1) (EDRR 2011: 0)]
59->	United States, Maryland, Keyser Ridge; 4/IV/2011; SC SW RB TD; trap alpha pinene + ethanol	[unknown (1) (EDRR 2011: 0)]
60->	United States, Maryland, Maryland (state)	[unknown () (Wood, S.L. 1982: 270)]
61->	United States, Maryland, Plummers Island	[USNM () (TH Atkinson-1988, unpublished)]
62->	United States, Michigan, Michigan (state)	[unknown () (Wood, S.L. 1982: 270)]
63->	United States, Michigan, Williamston; 1/V/2011; Cognato, A.I.; trap alpha pinene + ethanol	[unknown (1) (EDRR 2011: 0)]
64->	United States, Mississippi, Agricultural College	[USNM () (TH Atkinson-1988, unpublished)]
65->	United States, Mississippi, Mississippi (state)	[unknown () (Wood, S.L. 1982: 270)]
66->	United States, Mississippi, Ripley	[USNM () (TH Atkinson-1988, unpublished)]
67->	United States, Mississippi, Wallerville; Blackman, M.W.	[DFEC () (TH Atkinson-1988, unpublished)]
68->	United States, Missouri, Ashland; 9/IV/2008; Hook, B.; trap alpha pinene + ethanol	[unknown (1) (EDRR 2011: 0)]
69->	United States, Missouri, Columbia; 16/VI/2014; Doerhoff, R.; trap: Lindgren Funnel alpha pinene + ethanol	[unknown (1) (EDRR 2014: 0)]
70->	United States, Missouri, Columbia; 25/IV/2014; Doerhoff, R.; trap: Lindgren Funnel alpha pinene + ethanol	[unknown (1) (EDRR 2014: 0)]
71->	United States, Missouri, Concordia; 10/IV/2008; Hook, B.; trap alpha pinene + ethanol	[unknown (1) (EDRR 2011: 0)]
72->	United States, Missouri, Eminence	[USNM () (TH Atkinson-1988, unpublished)]
73->	United States, Missouri, Finley River, 20 mi SE Sprigfield; 24/VI/1981; Haswell, R.H.	[TAMU (1) (TH Atkinson-2012, unpublished)]
74->	United States, Missouri, Kansas City	[USNM () (TH Atkinson-1988, unpublished)]
75->	United States, Missouri, Kansas City; 22/IV/2008; Hook, B.; trap alpha pinene + ethanol	[unknown (1) (EDRR 2011: 0)]
76->	United States, Missouri, Missouri (state)	[unknown () (Wood, S.L. 1982: 270)]
77->	United States, Missouri, Platte City; 23/IV/2014; Doerhoff, R.; trap: Lindgren Funnel alpha pinene + ethanol	[unknown (1) (EDRR 2014: 0)]
78->	United States, Missouri, Remote location; 12/V/2014; Doerhoff, R.; trap: Lindgren Funnel exotic ips lure	[unknown (1) (EDRR 2014: 0)]
79->	United States, Missouri, St. Louis	[MCZ () (TH Atkinson-1988, unpublished)]
80->	United States, Missouri, St. Louis; 12/IV/2008; Hook, B.; trap alpha pinene + ethanol	[unknown (1) (EDRR 2011: 0)]
81->	United States, Missouri, Warrenton; 11/IV/2008; Hook, B.; trap alpha pinene + ethanol	[unknown (1) (EDRR 2011: 0)]
82->	United States, Missouri, Weldon Spring; 17/IV/2014; Doerhoff, R.; trap: Lindgren Funnel ethanol	[unknown (1) (EDRR 2014: 0)]
83->	United States, Nebraska, Lincoln; 14/IV/2008; Harrell, M.; trap alpha pinene + ethanol	[unknown (1) (EDRR 2011: 0)]
84->	United States, Nebraska, Niobrara; 8/IV/2008; Harrell, M.; trap alpha pinene + ethanol	[unknown (2) (EDRR 2011: 0)]
85->	United States, Nebraska, Plattsmouth; 15/IV/2008; Harrell, M.; trap alpha pinene + ethanol	[unknown (1) (EDRR 2011: 0)]
86->	United States, Nebraska, Valentine; 8/IV/2008; Harrell, M.; trap alpha pinene + ethanol	[unknown (1) (EDRR 2011: 0)]
87->	United States, New Jersey, Clementon	[MCZ () (TH Atkinson-1988, unpublished)]
88->	United States, New Jersey, Mercer Co.; 17/IV/1937; Scott, V.F.	[FSCA (1) (TH Atkinson-2014, unpublished)]
89->	United States, New Jersey, New Jersey (state)	[unknown () (Wood, S.L. 1982: 270)]
90->	United States, New Jersey, Newark	[MCZ () (TH Atkinson-1988, unpublished)]
91->	United States, New York, Bronx; 31/V/2014; Cancelliere, J.; Denham, J.; trap: Lindgren Funnel exotic ips lure	[unknown (1) (EDRR 2014: 0)]
92->	United States, New York, New York (state)	[unknown () (Wood, S.L. 1982: 270)]
93->	United States, New York, Red Hook	[USNM () (TH Atkinson-1988, unpublished)]
94->	United States, New York, West Point	[USNM () (TH Atkinson-1988, unpublished)]
95->	United States, North Carolina, Canton; 19/IV/2011; Lawing; Heath; trap alpha pinene + ethanol	[unknown (1) (EDRR 2011: 0)]
96->	United States, North Carolina, Durham; Moraceae: Morus rubra	[Unknown () (Beal, J.A., Massey, C.L. 1945: 764)]
97->	United States, North Carolina, Durham; Cannabaceae: Celtis occidentalis	[unknown () (Beal, J.A., Massey, C.L. 1945: 77)]
98->	United States, North Carolina, Lexington; 25/IV/2013; Lawing; Heath; trap: Lindgren Funnel ethanol	[unknown (1) (EDRR 2014: 0)]
99->	United States, North Carolina, McDowell Co.; 1977; Weber, B.C.; trap: window trap	[unknown (1) (Weber, B.C., MacPherson, J.E. 1991: 46)]
100->	United States, North Carolina, North Carolina (state)	[unknown () (Wood, S.L. 1982: 270)]
101->	United States, North Carolina, Raleigh; 12/IV/2011; Blaedow, R.; trap alpha pinene + ethanol	[unknown (1) (EDRR 2011: 0)]
102->	United States, North Carolina, Tryon; Moraceae: Morus sp.	[USNM () (TH Atkinson-1988, unpublished)]
103->	United States, North Carolina, Tryon; Ulmaceae: Ulmus sp.	[USNM () (TH Atkinson-1988, unpublished)]
104->	United States, Ohio, Columbus	[USNM () (TH Atkinson-1988, unpublished)]
105->	United States, Ohio, Ohio (state)	[unknown () (Wood, S.L. 1982: 270)]
106->	United States, Oklahoma, Latimer Co.; V/2002; Stephan, K.; trap: flight intercept	[TAMU (2) (TH Atkinson-2012, unpublished)]
107->	United States, Oklahoma, Latimer Co.; IV/2002; Stephan, K.; trap: flight intercept	[TAMU (2) (TH Atkinson-2012, unpublished)]
108->	United States, Oklahoma, Red Oak; III/1988; Stephan, K.	[FSCA (2) (TH Atkinson-2007, unpublished)]
109->	United States, Oklahoma, Red Oak; VII/1989; Stephan, K.	[FSCA (3) (TH Atkinson-2007, unpublished)]
110->	United States, Oklahoma, Red Oak, 5 mi w; V/1982; Stephan, K.	[TAMU (1) (TH Atkinson-2007, unpublished) (SL Wood-1985, unpublished)]
111->	United States, Oklahoma, Red Oak, 5 mi w; 4/IV/1977; Stephan, K.	[TAMU (1) (SL Wood-2007, unpublished) (TH Atkinson-1985, unpublished)]
112->	United States, Oklahoma, Red Oak, 5 mi w; 28/II/1977; Stephan, K.	[TAMU (6) (SL Wood-2007, unpublished) (TH Atkinson-1985, unpublished)] [FSCA (12) (TH Atkinson-2007, unpublished)]
113->	United States, Oklahoma, Red Oak, 5 mi w; 18/III/1977; Stephan, K.	[TAMU (3) (SL Wood-2007, unpublished) (TH Atkinson-1985, unpublished)]
114->	United States, Oklahoma, Red Oak, 5 mi w; 25/III/1977; Stephan, K.	[TAMU (1) (SL Wood-2007, unpublished) (TH Atkinson-1985, unpublished)]
115->	United States, Oklahoma, Red Oak, 5 mi w; V/1985; Stephan, K.	[TAMU (1) (SL Wood-2007, unpublished) (TH Atkinson-1985, unpublished)]
116->	United States, Oklahoma, Red Oak, 5 mi w; V/1983; Stephan, K.	[TAMU (2) (SL Wood-2007, unpublished) (TH Atkinson-1985, unpublished)]
117->	United States, Oklahoma, Red Oak, 5 mi w; VII/1983; Stephan, K.	[TAMU (2) (SL Wood-2007, unpublished) (TH Atkinson-1985, unpublished)]
118->	United States, Oklahoma, Red Oak, 5 mi w; VI/1983; Stephan, K.	[TAMU (2) (SL Wood-2007, unpublished) (TH Atkinson-1985, unpublished)]
119->	United States, Oklahoma, Red Oak, 5 mi w; IV/1983; Stephan, K.	[TAMU (2) (SL Wood-2007, unpublished) (TH Atkinson-1985, unpublished)]
120->	United States, Oklahoma, Red Oak, 5 mi w; V/1984; Stephan, K.	[TAMU (3) (SL Wood-2007, unpublished) (TH Atkinson-1985, unpublished)]
121->	United States, Oklahoma, Red Oak, 5 mi w; IV/1984; Stephan, K.	[TAMU (1) (SL Wood-2007, unpublished) (TH Atkinson-1985, unpublished)]
122->	United States, Oklahoma, Red Oak, 5 mi w; X/1984; Stephan, K.	[TAMU (2) (SL Wood-2007, unpublished) (TH Atkinson-1985, unpublished)]
123->	United States, Oklahoma, Red Oak, 5 mi w; 1977; Stephan, K.	[USNM (8) (Atkinson, T.H.; Rabaglia,R.J.; Peck,S.J.; Foltz,J.L. 1991: 156)]
124->	United States, Pennsylvania, Abington	[MCZ () (TH Atkinson-1988, unpublished)]
125->	United States, Pennsylvania, Castle Rock	[USNM () (TH Atkinson-1988, unpublished)]
126->	United States, Pennsylvania, Delaware Co.	[USNM () (TH Atkinson-1988, unpublished)]
127->	United States, Pennsylvania, Frankfort (Springs?)	[USNM () (TH Atkinson-1988, unpublished)]
128->	United States, Pennsylvania, Harrisburg; 2/IV/2009; Spichiger, S.; trap alpha pinene + ethanol	[unknown (1) (EDRR 2011: 0)]
129->	United States, Pennsylvania, Manheim; 20/IV/2011; Schmit, D.; trap alpha pinene + ethanol	[unknown (1) (EDRR 2011: 0)]
130->	United States, Pennsylvania, Pennsylvania (state)	[unknown () (Wood, S.L. 1982: 270)]
131->	United States, Pennsylvania, Westown; Leguminosae: Robinia sp.	[USNM () (TH Atkinson-1988, unpublished)]
132->	United States, South Carolina, Clemson; 19/III/1938; Cartwright, O.L.	[CUAC (1) (TH Atkinson-1988, unpublished)]
133->	United States, South Carolina, Clemson; 29/V/1964; Burley, B.	[CUAC (1) (TH Atkinson-1988, unpublished)]
134->	United States, South Carolina, Rock Hill; 17/III/2010; Reid, L.; trap alpha pinene + ethanol	[unknown (1) (EDRR 2011: 0)]
135->	United States, South Carolina, South Carolina (state)	[unknown () (Wood, S.L. 1982: 270)]
136->	United States, South Carolina, Spartanburg Co.; 21/V/1984; Kovach, J.S.	[CUAC (1) (TH Atkinson-1988, unpublished)]
137->	United States, Tennessee, Kingsport; 13/V/2009; Bingham, S.; trap alpha pinene + ethanol	[unknown (1) (EDRR 2011: 0)]
138->	United States, Tennessee, Knoxville; 13/V/2009; Hansard, J.; trap alpha pinene + ethanol	[unknown (2) (EDRR 2011: 0)]
139->	United States, Tennessee, Lawrenceburg; 25/VII/1951; Price, R.D.; Beamer, R.H.; Beamer, L.D.; Wood, S.L; Moraceae: Morus sp.	[USNM (2) (Wood, S.L. 1982: 270) (TH Atkinson-1987, unpublished)]
140->	United States, Tennessee, Memphis	[USNM () (TH Atkinson-1988, unpublished)]
141->	United States, Texas, Baltrane; Moraceae: Maclura pomifera	[USNM (2) (TH Atkinson-1988, unpublished)]
142->	United States, Texas, Columbus	[USNM () (TH Atkinson-1988, unpublished)]
143->	United States, Texas, Dallas; 31/III/1951; Wood, S.L.; Cannabaceae: Celtis laevigata	[USNM (16) (Wood, S.L. 1982: 270) (TH Atkinson-1988, unpublished)]
144->	United States, Texas, Devil's River	[USNM () (TH Atkinson-1988, unpublished)]
145->	United States, Texas, Elkhart, 10 mi SW; 24/XII/1964; Burke, H.R.	[TAMU (1) (TH Atkinson-, unpublished)]
146->	United States, Texas, Victoria	[USNM () (TH Atkinson-1988, unpublished)]
147->	United States, Texas, Welder Wildlife Ref; 20/III/1965; Chamberlin, W.F.	[TAMU (3) (TH Atkinson-2003, unpublished)]
148->	United States, Texas, Westlake Hills, Lake Austin at River Hills Drive; 29/XI/2012; Atkinson, T.H.; trap: alpha-scents 4 vane ethanol	[UTIC (4) (TH Atkinson-2013, unpublished)]
149->	United States, Texas, Woodville; 16/IV/2013; Atkinson, T.H.; trap: alpha-scents 4 vane ethanol	[UTIC (1) (TH Atkinson-2013, unpublished)]
150->	United States, Texas, Woodville, 5 mi E; 6/III/2013; Atkinson, T.H.; trap: alpha-scents 4 vane ethanol	[UTIC (1) (TH Atkinson-2013, unpublished)]
151->	United States, Virginia, Falls Church; Moraceae: Morus sp.	[USNM () (TH Atkinson-1988, unpublished)]
152->	United States, Virginia, Virginia (state)	[unknown () (Wood, S.L. 1982: 270)]
153->	United States, Virginia, Virginia Beach	[USNM () (TH Atkinson-1988, unpublished)]
154->	United States, West Virginia, Cairo	[USNM () (TH Atkinson-1988, unpublished)]
155->	United States, West Virginia, Kanawha Sta.; Moraceae: Morus rubra	[USNM () (TH Atkinson-1988, unpublished)]
156->	United States, West Virginia, Kanawha Sta.	[USNM () (TH Atkinson-1988, unpublished)]
157->	United States, West Virginia, Monongalia Co.	[USNM () (TH Atkinson-1988, unpublished)]
158->	United States, West Virginia, Morgantown	[USNM () (TH Atkinson-1988, unpublished)]
159->	United States, West Virginia, New Martinsville	[USNM () (TH Atkinson-1988, unpublished)]
160->	United States, West Virginia, Preston Co.	[USNM () (TH Atkinson-1988, unpublished)]
161->	United States, West Virginia, Wayne Co.	[USNM () (TH Atkinson-1988, unpublished)]
162->	United States, West Virginia, West Virginia (state)	[unknown () (Wood, S.L. 1982: 270)]
163->	United States, West Virginia, Wood Co.	[USNM () (TH Atkinson-1988, unpublished)]