Bark and Ambrosia Beetles of

SYNONYMY

• Dendroctonus frontalis Zimmermann 1868. Trans. Amer. Ent. Soc. 2: 149
• Dendroctonus arizonicus Hopkins 1909. U.S. Dep. Agric. Tech. Bull. 17(1): 95

DISTRIBUTION.

Numbers in parentheses after each geographic unit are the number of distinct collection events in the database for that unit. For exotic species generally only countries are listed for localities outside the New World. For further information on published sources of distribution, check the REFERENCES section.

Distribution Comments.   This species and D. mexicanus have been confused and considered synonyms in the past. As a consequence the actual distributions and degree of overlap of these 2 species in Arizona and Mexico is not entirely clear.

• Central America (21)
  Belize (1): Cayo (1); Guatemala (8): Guatemala (2), Huehuetenango (1), Sololá (3), unknown (1), Zacapa (1); Honduras (10): Atlántida (1), El Paraíso (2), Francisco Morazán (4), Olancho (1), Yoro (2); Nicaragua (2): Chinandega (1), Nueva Segovia (1);
• North America (144)
  (1): no state / prov. data (1); México (27): Chiapas (4), Chihuahua (1), Durango (1), Guerrero (1), Hidalgo (1), Jalisco (1), México (2), Michoacán (3), Morelos (1), Nuevo León (1), Oaxaca (6), Puebla (1), Querétaro (2), San Luís Potosí (1), Tamaulipas (1); United States (116): no state / prov. data (7), Alabama (2), Arizona (13), Arkansas (2), Connecticut (5), Delaware (1), District of Columbia (1), Florida (6), Georgia (12), Louisiana (7), Maryland (1), New York (1), North Carolina (12), Oklahoma (1), Pennsylvania (2), Rhode Island (1), South Carolina (8), Tennessee (1), Texas (10), Virginia (12), West Virginia (11);

HOSTS

Numbers in parentheses after each host family, genus, or collection method are the number of distinct collection events in the database for that host or method.

• No host information (108);
• Pinaceae (40): Pinus (40): no species data (1), caribaea (1), chihuahuana (3), clausa (1), echinata (3), engelmannii (1), glabra (1), greggii (1), oocarpa (7), palustris (1), ponderosa (3), pringlei (3), pseudostrobus (3), rigida (2), sp. (3), strobus (1), taeda (4), virginiana (1);
• trap (17) (5): alpha pinene + ethanol (5); light (3): mercury vapor (1), mercury vapor + UV (2); Lindgren Funnel (9): no details (2), alpha pinene + ethanol (3), exotic ips lure (2), frontalin + alpha pinene (2);

REFERENCES

The following are important recent monographs, catalogs, and supplements to catalogs that refer to this species. The specific page on which the reference is made is shown in pink at the end of the reference. In the case of Wood (1982) and Wood & Bright (1992) clicking on the reference page will link to a digital version of the work in question.

• Wood, S.L. 1982. The bark and ambrosia beetles of North and Central America (Coleoptera: Scolytidae), a taxonomic monograph. Great Basin Nat. Mem. 6:1-1356. [159]. (data capture complete)
• Wood, S.L., Bright,D.E. 1992. A catalog of Scolytidae and Platypodidae (Coleoptera), Part 2. Taxonomic Index. Great Basin Nat. Mem. 13:1-1553 (vol. A, B). [143]

The following are references from which host and distribution data have been input into the database. If one of the above monographs or catalogs also appears in this list, it means that most relevant collection event data have been included.

1. . . .
2. Anonymous, 2010. Michigan State University. SCAN: Symbiota Collections of Artropods Network. http://symbiota4.acis.ufl.edu/scan/portal/collections/misc/collprofiles.php?collid=76: -.
3. Armendariz, F.; Niño, A.; Sullivan, B.T.; Macías Samaño, J.; Victor, J.; Clarke, S.R.; Zúñiga, G. 2014. Two Species Within Dendroctonus frontalis (Coleoptera: Curculionidae): Evidence From Morphological, Karyological, Molecular, and Crossing Studies. Ann. Entomol. Soc. Am. 107(1): 11-27.
4. Bright, D.E., Skidmore, R.E. 2002. A Catalog of Scolytidae and Platypodidae (Coleoptera), Supplement 2 (1995-1999). NRC Research Press. Ottawa, Ontario, Canada.
5. EDRR 2011. USFS Early Detection Rapid Response Database. http://www.fs.fed.us/invasivespecies/earlydetection.shtml/.
6. EDRR 2014. USFS Early Detection Rapid Response Database. http://www.fs.fed.us/invasivespecies/earlydetection.shtml/.
7. Moser, J.C.; Fitzgibbon, B.A.; Klepzig, K.D. 2005. The Mexican pine beetle, Dendroctonus mexicanus : first record in the United States and co-occurrence with the southern pine beetle, Dendroctonus frontalis (Coleoptera: Scolytidae or Curculionidae:Scolytinae). Entomological News 116 (4): 235-243.
8. Rabaglia, R.J.; Valenti, M.A. 2003. Annotated list of the bark and ambrosia beetles (Coleoptera: Scolytidae) of Delaware, with new distribution records. Proc. Entomol. Soc. Wash. 105(2): 312-319.
9. Salinas Moreno, Y.; Zúñiga, G.; Cisneros, R. 1994. Chromosomal variation in Mexican populations of the genus <i>Dendroctonus</i> (Coleoptera: Scolytidae). . SW Natur. 39: 283-286.
10. University of Arizona Insect Collection 2016. SCAN: Symbiota Collections of Arthropods Network. http://symbiota4.acis.ufl.edu/scan/portal/collections/misc/collprofiles.php?collid=11.
11. Wood, S.L. 1982. The bark and ambrosia beetles of North and Central America (Coleoptera: Scolytidae), a taxonomic monograph. Great Basin Nat. Mem. 6: 1-1356.
12. Wood, S.L.; Bright, D.E. 1992. A catalog of Scolytidae and Platypodidae (Coleoptera), Part 2. Taxonomic Index (Volumes A,B). Great Basin Nat. Mem. 13: 1-1553.
13. Zúñiga, G.; Cisneros, R.; Hayes, J.L.; Macías Samaño, J. 2002. Karyology, Geographic Distribution, and Origin of the Genus Dendroctonus Erichson (Coleoptera: Scolytidae). Ann. Entomol. Soc. Am. 95(3): 267-275.
14. Zúñiga, G.; Cisneros, R.; Salinas Moreno, Y. 1995. Coexistencia de Dendroctonus frontalis Zimmermann y D. mexicanus Hopkins (Coleoptera: Scolytidae) sobre un mismo hospedero. Acta Zool. Mex. N. S. 64: 59-62.
15. Zúñiga, G.; Cisneros, R.; Salinas Moreno, Y. 1998. Chromosome numbers of Dendroctonus parallelocollis and D. rhizophagus ( Coleoptera: Scolytidae) and new reports of Dendroctonus populations from Mexico and Guatemala. Ann. Entomol. Soc. Am. 91(4): 392-394.

Maps automatically open at the center of the plotted points and the scale is set to encompass all map points. Maps can be resized (scale bar at upper left) and the center moved (place cursor over map and drag) to see other parts of the distribution of the species. Clicking on a map marker will pull up collection event data and a literature citation if present. If any errors are found, please refer to the "series code" which is a unique identifier for a database record in any communications). Coordinates have not been entered for all collection records. Localities outside the New World are not plotted, even though they are listed in the distribution summary and in the table of records.

No.	Collection data	Publication
Central America
1->	Belize, Cayo, Mountain pine ridge; trap: Lindgren Funnel	[unknown (1) (Armendariz, F.; Niño, A.; Sullivan, B.T.; Macías Samaño, J.; Victor, J.; Clarke, S.R.; Zúñiga, G. 2014: 14)]
2->	Guatemala, Guatemala, Amatitlán	[unknown (1) (Zúñiga, G.; Cisneros, R.; Salinas Moreno, Y. 1998: 394) (Zúñiga, G.; Cisneros, R.; Hayes, J.L.; Macías Samaño, J. 2002: 274)]
3->	Guatemala, Guatemala, San Pedro Ayampuc	[unknown (1) (Zúñiga, G.; Cisneros, R.; Salinas Moreno, Y. 1998: 394) (Zúñiga, G.; Cisneros, R.; Hayes, J.L.; Macías Samaño, J. 2002: 274)]
4->	Guatemala, Huehuetenango, Caselaj; Pinaceae: Pinus oocarpa	[unknown (1) (Armendariz, F.; Niño, A.; Sullivan, B.T.; Macías Samaño, J.; Victor, J.; Clarke, S.R.; Zúñiga, G. 2014: 14)]
5->	Guatemala, Sololá, Colonia María Tecun; Pinaceae: Pinus oocarpa	[unknown (1) (Armendariz, F.; Niño, A.; Sullivan, B.T.; Macías Samaño, J.; Victor, J.; Clarke, S.R.; Zúñiga, G. 2014: 14)]
6->	Guatemala, Sololá, Finca Chuchiya; Pinaceae: Pinus pseudostrobus	[unknown (1) (Armendariz, F.; Niño, A.; Sullivan, B.T.; Macías Samaño, J.; Victor, J.; Clarke, S.R.; Zúñiga, G. 2014: 14)]
7->	Guatemala, Sololá, Finca Socorro; Pinaceae: Pinus pseudostrobus	[unknown (1) (Armendariz, F.; Niño, A.; Sullivan, B.T.; Macías Samaño, J.; Victor, J.; Clarke, S.R.; Zúñiga, G. 2014: 14)]
8->	Guatemala, unknown, Guatemala (country)	[unknown (1) (Wood, S.L.; Bright, D.E. 1992: 143)]
9->	Guatemala, Zacapa, Usumatlán	[unknown (1) (Zúñiga, G.; Cisneros, R.; Salinas Moreno, Y. 1998: 394) (Zúñiga, G.; Cisneros, R.; Hayes, J.L.; Macías Samaño, J. 2002: 274)]
10->	Honduras, Atlántida, Parque Nal. Pico Bonito, Est. CURLA; 6/VI/2003; Turnbow, R.H.; Pinaceae: Pinus	[RHTC (1) (TH Atkinson-2008, unpublished)]
11->	Honduras, El Paraíso, Yuscarán	[unknown () (Wood, S.L. 1982: 159)]
12->	Honduras, El Paraíso, Yuscaran; 21/IV/1964; Wood, S.L.	[MSUC (1) (Anonymous, 2010: 0)]
13->	Honduras, Francisco Morazán, Cerro Uyuca; 15/VII/2001; Turnbow, R.H.; trap: light mercury vapor + UV	[RHTC (1) (TH Atkinson-2008, unpublished)]
14->	Honduras, Francisco Morazán, Guaimaca; Pinaceae: Pinus oocarpa	[unknown (1) (Armendariz, F.; Niño, A.; Sullivan, B.T.; Macías Samaño, J.; Victor, J.; Clarke, S.R.; Zúñiga, G. 2014: 14)]
15->	Honduras, Francisco Morazán, Tegucigalpa; 25/II/1966; Wilkinson, R.C.; Pinaceae: Pinus oocarpa	[FSCA (2) (TH Atkinson-1988, unpublished)]
16->	Honduras, Francisco Morazán, Tegucigalpa; 1/III/1966; Wilkinson, R.C.; Pinaceae: Pinus oocarpa	[FSCA (1) (TH Atkinson-2007, unpublished)]
17->	Honduras, Olancho, Olancho; 7/I/1964; Banegas, A.D.	[TAMU (8) (TH Atkinson-, unpublished)]
18->	Honduras, Yoro, Las Cumbas; 14/V/2002; Turnbow, R.H.; trap: light mercury vapor	[RHTC (4) (TH Atkinson-2008, unpublished)]
19->	Honduras, Yoro, Parque Nal. Pico Pijol; 22/VII/2001; Turnbow, R.H.; trap: light mercury vapor + UV	[RHTC (5) (TH Atkinson-2008, unpublished)]
20->	Nicaragua, Chinandega, El Picacho, Cerro Tomabu; Pinaceae: Pinus oocarpa	[unknown (1) (Armendariz, F.; Niño, A.; Sullivan, B.T.; Macías Samaño, J.; Victor, J.; Clarke, S.R.; Zúñiga, G. 2014: 14)]
21->	Nicaragua, Nueva Segovia, Jalapa; Pinaceae: Pinus caribaea	[unknown (1) (Armendariz, F.; Niño, A.; Sullivan, B.T.; Macías Samaño, J.; Victor, J.; Clarke, S.R.; Zúñiga, G. 2014: 14)]
North America
22->	, no state / prov. data, no label data	[UAIC (1) (University of Arizona Insect Collection 2016: 0)]
23->	México, Chiapas, Buenavista	[unknown (1) (Zúñiga, G.; Cisneros, R.; Hayes, J.L.; Macías Samaño, J. 2002: 274) (Armendariz, F.; Niño, A.; Sullivan, B.T.; Macías Samaño, J.; Victor, J.; Clarke, S.R.; Zúñiga, G. 2014: 14)]
24->	México, Chiapas, Chiapas (estado)	[unknown (1) (Wood, S.L.; Bright, D.E. 1992: 143)]
25->	México, Chiapas, Dos Lagunas	[unknown (1) (Zúñiga, G.; Cisneros, R.; Hayes, J.L.; Macías Samaño, J. 2002: 274) (Armendariz, F.; Niño, A.; Sullivan, B.T.; Macías Samaño, J.; Victor, J.; Clarke, S.R.; Zúñiga, G. 2014: 14)]
26->	México, Chiapas, La Trinitaria, Parque Nacional Lagunas de Montebello; Pinaceae: Pinus oocarpa	[unknown (1) (Armendariz, F.; Niño, A.; Sullivan, B.T.; Macías Samaño, J.; Victor, J.; Clarke, S.R.; Zúñiga, G. 2014: 14)]
27->	México, Chihuahua, Chihuahua (estado)	[unknown (1) ( : 12)]
28->	México, Durango, Durango (estado)	[unknown (0) (Bright, D.E., Skidmore, R.E. 2002: 26)]
29->	México, Guerrero, Guerrero (estado)	[unknown (1) (Wood, S.L.; Bright, D.E. 1992: 143)]
30->	México, Hidalgo, Hidalgo (estado)	[unknown (1) ( : 12)]
31->	México, Jalisco, Jalisco (estado)	[unknown (1) ( : 12)]
32->	México, México, Cascadas de Nanchititla, Paraje Los Venados	[unknown (1) (Salinas Moreno, Y.; Zúñiga, G.; Cisneros, R. 1994: 286) (Zúñiga, G.; Cisneros, R.; Hayes, J.L.; Macías Samaño, J. 2002: 274)]
33->	México, México, México (estado)	[unknown (0) (Bright, D.E., Skidmore, R.E. 2002: 26)]
34->	México, Michoacán, La Mesa	[unknown (1) (Salinas Moreno, Y.; Zúñiga, G.; Cisneros, R. 1994: 286) (Zúñiga, G.; Cisneros, R.; Hayes, J.L.; Macías Samaño, J. 2002: 274)]
35->	México, Michoacán, Michoacán (state)	[unknown (0) (Bright, D.E., Skidmore, R.E. 2002: 26)]
36->	México, Michoacán, Ojo de Agua de los Sauces	[unknown (1) (Salinas Moreno, Y.; Zúñiga, G.; Cisneros, R. 1994: 286) (Zúñiga, G.; Cisneros, R.; Hayes, J.L.; Macías Samaño, J. 2002: 274)]
37->	México, Morelos, Morelos (estado)	[unknown (1) ( : 12)]
38->	México, Nuevo León, Nuevo León (estado)	[unknown (1) ( : 12)]
39->	México, Oaxaca, 'Oaxaca norte', coordinates; Pinaceae: Pinus pringlei	[unknown (1) (Armendariz, F.; Niño, A.; Sullivan, B.T.; Macías Samaño, J.; Victor, J.; Clarke, S.R.; Zúñiga, G. 2014: 14)]
40->	México, Oaxaca, Los Azufres	[unknown (1) (Zúñiga, G.; Cisneros, R.; Hayes, J.L.; Macías Samaño, J. 2002: 274) (Salinas Moreno, Y.; Zúñiga, G.; Cisneros, R. 1994: 286)]
41->	México, Oaxaca, Oaxaca (estado)	[unknown (0) (Bright, D.E., Skidmore, R.E. 2002: 26)]
42->	México, Oaxaca, San Pedro Ayutla, Zacatepec; Pinaceae: Pinus pringlei	[unknown (1) (Armendariz, F.; Niño, A.; Sullivan, B.T.; Macías Samaño, J.; Victor, J.; Clarke, S.R.; Zúñiga, G. 2014: 14)]
43->	México, Oaxaca, Santa Cruz Nundaco	[unknown (1) (Zúñiga, G.; Cisneros, R.; Salinas Moreno, Y. 1998: 394) (Zúñiga, G.; Cisneros, R.; Hayes, J.L.; Macías Samaño, J. 2002: 274)]
44->	México, Oaxaca, Sta. Ma. Albarradas, San Pablo Villa de Mitla; Pinaceae: Pinus pringlei	[unknown (1) (Armendariz, F.; Niño, A.; Sullivan, B.T.; Macías Samaño, J.; Victor, J.; Clarke, S.R.; Zúñiga, G. 2014: 14)]
45->	México, Puebla, Puebla (estado)	[unknown (1) ( : 12)]
46->	México, Querétaro, La Parada; Pinaceae: Pinus greggii	[unknown (1) (Armendariz, F.; Niño, A.; Sullivan, B.T.; Macías Samaño, J.; Victor, J.; Clarke, S.R.; Zúñiga, G. 2014: 14)]
47->	México, Querétaro, Los Herrera; II/1993; Zúñiga, G.; Cisneros, R.; Salinas, Y.; Pinaceae: Pinus pseudostrobus	[unknown (1) (Zúñiga, G.; Cisneros, R.; Salinas Moreno, Y. 1995: 59) (Salinas Moreno, Y.; Zúñiga, G.; Cisneros, R. 1994: 286) (Zúñiga, G.; Cisneros, R.; Hayes, J.L.; Macías Samaño, J. 2002: 274)]
48->	México, San Luís Potosí, San Luís Potosí (estado)	[unknown (1) (TH Atkinson-2017, unpublished)]
49->	México, Tamaulipas, Tamaulipas (estado)	[unknown (1) ( : 12)]
50->	United States, no state / prov. data; Pinaceae: Pinus engelmannii	[unknown (0) (Wood, S.L. 1982: 159)]
51->	United States, no state / prov. data; Pinaceae: Pinus glabra	[unknown (0) (Wood, S.L. 1982: 159)]
52->	United States, no state / prov. data; Pinaceae: Pinus palustris	[unknown (0) (Wood, S.L. 1982: 159)]
53->	United States, no state / prov. data; Pinaceae: Pinus ponderosa	[unknown (0) (Wood, S.L. 1982: 159)]
54->	United States, no state / prov. data; Pinaceae: Pinus rigida	[unknown (0) (Wood, S.L. 1982: 159)]
55->	United States, no state / prov. data; Pinaceae: Pinus strobus	[unknown (0) (Wood, S.L. 1982: 159)]
56->	United States, no state / prov. data, Carolina	[MCZ (1: holotype, male)
57->	United States, Alabama, Calhoun	[unknown () (Wood, S.L. 1982: 159)]
58->	United States, Alabama, Montgomery	[unknown () (Wood, S.L. 1982: 159)]
59->	United States, Arizona, Brush Corral; 16/XII/1914; Edmonston, W.D.; Pinaceae: Pinus chihuahuana	[UAIC (2) (University of Arizona Insect Collection 2016: 0)]
60->	United States, Arizona, Flagstaff; Webb, J.L.; Pinaceae: Pinus ponderosa	[UAIC (1) (University of Arizona Insect Collection 2016: 0)]
61->	United States, Arizona, Pinery Canyon; 1/V/2001; Moser, J.C.; Fitzgibbon, B.A.; Klepzig, K.D.; trap: Lindgren Funnel frontalin + alpha pinene	[USFSP (1) (Moser, J.C.; Fitzgibbon, B.A.; Klepzig, K.D. 2005: 235)]
62->	United States, Arizona, Pinery Canyon; 6/XII/2001; Moser, J.C.; Fitzgibbon, B.A.; Klepzig, K.D.; Pinaceae: Pinus chihuahuana	[USFSP (1) (Moser, J.C.; Fitzgibbon, B.A.; Klepzig, K.D. 2005: 235)]
63->	United States, Arizona, Prescott, Copper Basin Road	[unknown () (Wood, S.L. 1982: 159)]
64->	United States, Arizona, San Francisco Mtn., Schultz Pass Road; 16/VIII/1998; Baranowski, R.M.; Pinaceae: Pinus ponderosa	[ZMLU (1) (TH Atkinson-2017, unpublished)]
65->	United States, Arizona, Sta. Catalina Mtns.	[unknown () (Wood, S.L. 1982: 159)]
66->	United States, Arizona, Sta. Catalina Mtns.; 5/IV/1969	[FSCA (1) (DE Bright-2010, unpublished) (TH Atkinson-2013, unpublished)]
67->	United States, Arizona, Sta. Catalina Mtns., Mt. Lemmon; 11/VI/1984; Baranowski, R.M.	[ZMLU (1) (TH Atkinson-2017, unpublished)]
68->	United States, Arizona, Turkey Creek; 4/III/2002; Moser, J.C.; Fitzgibbon, B.A.; Klepzig, K.D.; trap: Lindgren Funnel frontalin + alpha pinene	[USFSP (1) (Moser, J.C.; Fitzgibbon, B.A.; Klepzig, K.D. 2005: 235)]
69->	United States, Arizona, Turkey Creek; 3/X/2001; Moser, J.C.; Fitzgibbon, B.A.; Klepzig, K.D.; Pinaceae: Pinus chihuahuana	[USFSP (1) (Moser, J.C.; Fitzgibbon, B.A.; Klepzig, K.D. 2005: 235)]
70->	United States, Arizona, Walker	[unknown () (Wood, S.L. 1982: 159)]
71->	United States, Arizona, Williams; 6/VII/1901; Barber, H.S.; Schwarz, E.A.	[USNM (1: holotype, female) (Wood, S.L. 1982: 159) (TH Atkinson-2018, unpublished)]
72->	United States, Arkansas, Hampton	[unknown () (Wood, S.L. 1982: 159)]
73->	United States, Arkansas, Oachita Natl. For.; 31/III/1983; Barr, C.B.; Barr, J.E.; Pinaceae: Pinus sp.	[LSAM (1) (TH Atkinson-1988, unpublished)]
74->	United States, Connecticut, Hartford Co.; 2015; NAPIS Survey	[unknown (0) (unknown-2015, unpublished)]
75->	United States, Connecticut, Litchfield Co.; 2015; NAPIS Survey	[unknown (0) (unknown-2015, unpublished)]
76->	United States, Connecticut, Middlesex Co.; 2015; NAPIS Survey	[unknown (0) (unknown-2015, unpublished)]
77->	United States, Connecticut, New Haven Co.; 2015; NAPIS Survey	[unknown (0) (unknown-2015, unpublished)]
78->	United States, Connecticut, New London Co.; 2015; NAPIS Survey	[unknown (0) (unknown-2015, unpublished)]
79->	United States, Delaware, Oak Orchard; 5/I/1961; Bray, D.F.; Pinaceae: Pinus taeda	[UDCC (9) (Rabaglia, R.J.; Valenti, M.A. 2003: 314)]
80->	United States, District of Columbia, Washington	[unknown () (Wood, S.L. 1982: 159)]
81->	United States, Florida, Haw Creek	[Unknown () (-, unpublished)]
82->	United States, Florida, Haw Creek	[unknown () (Wood, S.L. 1982: 159)]
83->	United States, Florida, Marion Co.; 12/XII/1947; Capelouto, R.; Pinaceae: Pinus taeda	[FSCA (7) (TH Atkinson-2013, unpublished)]
84->	United States, Florida, Tallahassee; 12/X/1974; Wilkinson, R.C.; Pinaceae: Pinus echinata	[TAMU (99) (TH Atkinson-2005, unpublished)]
85->	United States, Florida, Taylor Ck	[Unknown () (-, unpublished)]
86->	United States, Florida, Walton Co.; 29/X/1975; Atkinson, T.H.; Pinaceae: Pinus clausa	[TAMU (1) (TH Atkinson-1989, unpublished)]
87->	United States, Georgia, Athens; 20/XI/1963; Nelms, M.	[FSCA (1) (TH Atkinson-2013, unpublished) (DE Bright-2010, unpublished)]
88->	United States, Georgia, Clarke Co.; 13/XI/1975; Gargiullo, P.	[UGCA (1) (TH Atkinson-1988, unpublished)]
89->	United States, Georgia, Clyo	[unknown () (Wood, S.L. 1982: 159)]
90->	United States, Georgia, Commerce; 10/VII/1988; Moreira, A.; Pinaceae: Pinus taeda	[A. Moreira (1) (TH Atkinson-1988, unpublished)]
91->	United States, Georgia, Commerce; 14/VII/1988; Moreira, A.; Pinaceae: Pinus taeda	[A. Moreira (2) (TH Atkinson-1988, unpublished)]
92->	United States, Georgia, Demorest	[unknown () (Wood, S.L. 1982: 159)]
93->	United States, Georgia, Eden; 6/V/2014; Morgan, R.; trap: Lindgren Funnel alpha pinene + ethanol	[unknown (1) (EDRR 2014: 0)]
94->	United States, Georgia, Habersham Co; 14/XI/1962; Hazard, E.; Pinaceae: Pinus echinata	[FSCA (33) (TH Atkinson-1988, unpublished)]
95->	United States, Georgia, Jackson Co.; 23/II/1976; Hurd, D.A.; Pinaceae: Pinus sp.	[UGCA (2) (TH Atkinson-1988, unpublished)]
96->	United States, Georgia, Red Fox Run; VI/1975; Berisford, C.W.	[UGCA (15) (TH Atkinson-1988, unpublished)]
97->	United States, Georgia, Savannah; 7/V/2014; Morgan, R.; trap: Lindgren Funnel alpha pinene + ethanol	[unknown (1) (EDRR 2014: 0)]
98->	United States, Georgia, Thomasville	[unknown () (Wood, S.L. 1982: 159)]
99->	United States, Louisiana, Baton Rouge; 17/III/1985; Riley, E.G.	[EGRC (1) (TH Atkinson-1987, unpublished)]
100->	United States, Louisiana, Baton Rouge; 2/V/1982	[LSAM (1) (TH Atkinson-1988, unpublished)]
101->	United States, Louisiana, Baton Rouge; 15/IV/1986; Rider, D.A.	[LSAM (4) (TH Atkinson-1988, unpublished)]
102->	United States, Louisiana, Hammond; 15/III/2007; Quebedeaux, J.; trap alpha pinene + ethanol	[unknown (1) (EDRR 2011: 0)]
103->	United States, Louisiana, Idlewild Exp Sta; 21/X/1982; McAuley, M.A.	[TAMU (24) (TH Atkinson-2011, unpublished)]
104->	United States, Louisiana, Singer	[unknown () (Wood, S.L. 1982: 159)]
105->	United States, Louisiana, Wilson	[unknown () (Wood, S.L. 1982: 159)]
106->	United States, Maryland, Cumberland	[unknown () (Wood, S.L. 1982: 159)]
107->	United States, New York, Wertheim NWR; 14/X/2014; Gilrein, D,; Pinaceae: Pinus rigida	[UTIC (3) (TH Atkinson-2021, unpublished)]
108->	United States, North Carolina, Asheville; VII/1959; Merkle, E.; Pinaceae: Pinus echinata	[FSCA (14) (TH Atkinson-1988, unpublished)]
109->	United States, North Carolina, Asheville; VII/1959; Merkle, E.; Pinaceae: Pinus virginiana	[FSCA (5) (TH Atkinson-1988, unpublished)]
110->	United States, North Carolina, Asheville	[unknown () (Wood, S.L. 1982: 159)]
111->	United States, North Carolina, Biltmore	[unknown () (Wood, S.L. 1982: 159)]
112->	United States, North Carolina, Black Mtn.	[unknown () (Wood, S.L. 1982: 159)]
113->	United States, North Carolina, Blue Ridge Pkwy; 19/III/1975; Turnbow, R.H.	[UGCA (3) (TH Atkinson-1988, unpublished)]
114->	United States, North Carolina, Boardman	[unknown () (Wood, S.L. 1982: 159)]
115->	United States, North Carolina, Broad Creek; 2/IV/2008; Blaedow, R.; trap alpha pinene + ethanol	[unknown (1) (EDRR 2011: 0)]
116->	United States, North Carolina, Fletcher	[unknown () (Wood, S.L. 1982: 159)]
117->	United States, North Carolina, Greybeard Mtn.	[unknown () (Wood, S.L. 1982: 159)]
118->	United States, North Carolina, Pisgah Ridge	[unknown () (Wood, S.L. 1982: 159)]
119->	United States, North Carolina, Tryon	[unknown () (Wood, S.L. 1982: 159)]
120->	United States, Oklahoma, Western Indian Terr.	[unknown () (Wood, S.L. 1982: 159)]
121->	United States, Pennsylvania, Mont Alto	[unknown () (Wood, S.L. 1982: 159)]
122->	United States, Pennsylvania, Nottingham; 13/VI/2017; Peters, G.; Smith, P.; trap: Lindgren Funnel	[UTIC (3) (TH Atkinson-2021, unpublished)]
123->	United States, Rhode Island, Washington Co.; 2015; NAPIS Survey	[unknown (0) (unknown-2015, unpublished)]
124->	United States, South Carolina, Ben Quan?	[unknown () (Wood, S.L. 1982: 159)]
125->	United States, South Carolina, Clemson	[unknown () (Wood, S.L. 1982: 159)]
126->	United States, South Carolina, Columbia; 2/IV/2012; Reid, L.; trap: Lindgren Funnel exotic ips lure	[unknown (1) (EDRR 2014: 0)]
127->	United States, South Carolina, Georgetown	[unknown () (Wood, S.L. 1982: 159)]
128->	United States, South Carolina, Ladson; 9/IV/2012; Reid, L.; trap: Lindgren Funnel exotic ips lure	[unknown (1) (EDRR 2014: 0)]
129->	United States, South Carolina, Laurens; 31/III/2007; Beckham, C.; trap alpha pinene + ethanol	[unknown (1) (EDRR 2011: 0)]
130->	United States, South Carolina, Modoc, 5.7 mi NE; 9/VII/1976; Turnbow, R.H.	[UGCA (1) (TH Atkinson-1988, unpublished)]
131->	United States, South Carolina, Pregnall	[unknown () (Wood, S.L. 1982: 159)]
132->	United States, Tennessee, Ducktown	[unknown () (Wood, S.L. 1982: 159)]
133->	United States, Texas, Bastrop; 17/III/2007; McAleese, K.; trap alpha pinene + ethanol	[unknown (1) (EDRR 2011: 0)]
134->	United States, Texas, Beaumont	[unknown () (Wood, S.L. 1982: 159)]
135->	United States, Texas, BTI Exp. For.; 15/I/1966; Guamer, G.C.	[TAMU (9) (TH Atkinson-2011, unpublished)]
136->	United States, Texas, Call	[unknown () (Wood, S.L. 1982: 159)]
137->	United States, Texas, Coldspring; VI/1983; Flamm, R.	[TAMU (42) (TH Atkinson-, unpublished)]
138->	United States, Texas, Deweyville	[unknown () (Wood, S.L. 1982: 159)]
139->	United States, Texas, Hardin Co.; 1958; Thatcher, R.C.	[TAMU (7) (TH Atkinson-2011, unpublished)]
140->	United States, Texas, Kirbyville	[unknown () (Wood, S.L. 1982: 159)]
141->	United States, Texas, Kirbyville, Seicke State Forest; 19/I/2007	[MSUC (1) (Anonymous, 2010: 0)]
142->	United States, Texas, Montgomery Co.; 19/VII/1962; Young; Pinaceae: Pinus sp.	[TAMU (10) (TH Atkinson-, unpublished)]
143->	United States, Virginia, Arlington	[unknown () (Wood, S.L. 1982: 159)]
144->	United States, Virginia, Auburn	[unknown () (Wood, S.L. 1982: 159)]
145->	United States, Virginia, Chase City	[unknown () (Wood, S.L. 1982: 159)]
146->	United States, Virginia, Cobb Isl	[unknown () (Wood, S.L. 1982: 159)]
147->	United States, Virginia, Glen Allen	[unknown () (Wood, S.L. 1982: 159)]
148->	United States, Virginia, Green Bay	[unknown () (Wood, S.L. 1982: 159)]
149->	United States, Virginia, King & Queen Co.	[unknown () (Wood, S.L. 1982: 159)]
150->	United States, Virginia, Newport News; 28/III/2008; Johnson; Ansicar; trap alpha pinene + ethanol	[unknown (1) (EDRR 2011: 0)]
151->	United States, Virginia, Port Republic	[unknown () (Wood, S.L. 1982: 159)]
152->	United States, Virginia, Sandston; 5/IV/2013; Hubbard, H.G.; trap: Lindgren Funnel alpha pinene + ethanol	[unknown (1) (EDRR 2014: 0)]
153->	United States, Virginia, Virginia Beach	[unknown () (Wood, S.L. 1982: 159)]
154->	United States, Virginia, York Co; 17/VIII/1943; Hetrick, L.A.	[FSCA (1) (TH Atkinson-1988, unpublished)]
155->	United States, West Virginia, Greenbrier	[unknown () (Wood, S.L. 1982: 159)]
156->	United States, West Virginia, Hampshire Co.	[unknown () (Wood, S.L. 1982: 159)]
157->	United States, West Virginia, Hardy Co.	[unknown () (Wood, S.L. 1982: 159)]
158->	United States, West Virginia, Kanawha Co.	[unknown () (Wood, S.L. 1982: 159)]
159->	United States, West Virginia, Monongalia Co.	[unknown () (Wood, S.L. 1982: 159)]
160->	United States, West Virginia, Pendleton (Co.?)	[unknown () (Wood, S.L. 1982: 159)]
161->	United States, West Virginia, Pocahontas Co.	[unknown () (Wood, S.L. 1982: 159)]
162->	United States, West Virginia, Raleigh Co.	[unknown () (Wood, S.L. 1982: 159)]
163->	United States, West Virginia, Randolph Co.	[unknown () (Wood, S.L. 1982: 159)]
164->	United States, West Virginia, Tucker Co.	[unknown () (Wood, S.L. 1982: 159)]
165->	United States, West Virginia, Wood Co.	[unknown () (Wood, S.L. 1982: 159)]