Bark and Ambrosia Beetles of

SYNONYMY

Taxonomic Comments.   Mercado & Negron (2014: 311) erected a new genus for this species, removing it from Hylurgops on purely morphological grounds. I question whether this is justified, as no phylogenetic analysis was provided. This species is readily distinguished from North American species of Hylurgops.

• Hylastes cristatus Mannerheim 1853. Bull.Soc. Imper. Nat. Moscou 25: 239
• Hylurgops subcostulatus (Mannerheim 1853) Bull.Soc. Imper. Nat. Moscou 25: 239
• Pachysquamus subcostulatus (Mannerheim 1853) Bull.Soc. Imper. Nat. Moscou 25: 239
• Hylurgops subcostulatus subcostulatus (Mannerheim 1853) Bull.Soc. Imper. Nat. Moscou 25: 239
• Hylastes subcostulatus Mannerheim 1853. Bull.Soc. Imper. Nat. Moscou 25: 239
• Hylastes alternans Chapuis 1869. Synopsis des Scolytides: 22
• Hylurgops subcostulatus alternans (Chapuis 1869) Synopsis des Scolytides: 22

DISTRIBUTION.

Numbers in parentheses after each geographic unit are the number of distinct collection events in the database for that unit. For exotic species generally only countries are listed for localities outside the New World. For further information on published sources of distribution, check the REFERENCES section.

Distribution Comments.   Western North America from southwestern Canada to Chiapas.

• North America (233)
  (3): no state / prov. data (3); Canada (7): British Columbia (7); México (44): no state / prov. data (1), Baja California (3), Chiapas (4), Chihuahua (3), Distrito Federal (1), Durango (3), Guerrero (2), Hidalgo (1), México (12), Michoacán (1), Morelos (5), Oaxaca (2), Puebla (2), unknown (1), Veracruz (3); United States (179): no state / prov. data (1), Alaska (2), Arizona (37), California (52), Colorado (28), Idaho (11), Montana (4), Nevada (2), New Mexico (5), Oregon (16), South Dakota (10), Utah (2), Washington (8), Wyoming (1);

HOSTS

Numbers in parentheses after each host family, genus, or collection method are the number of distinct collection events in the database for that host or method.

• No host information (147);
• Pinaceae (47): Picea (2): engelmanni (1), sp. (1); Pinus (44): no species data (2), ayacahuite (1), chihuahuana (1), hartwegii (4), jeffreyi (2), leiophylla (1), montezumae (3), ponderosa (11), sp. (19); Pseudotsuga (1): menziesii (1);
• trap (39) (25): alpha pinene + ethanol (25); Lindgren Funnel (13): alpha pinene + ethanol (5), ethanol (7), exotic ips lure (1); malaise (1): no details (1);

REFERENCES

The following are important recent monographs, catalogs, and supplements to catalogs that refer to this species. The specific page on which the reference is made is shown in pink at the end of the reference. In the case of Wood (1982) and Wood & Bright (1992) clicking on the reference page will link to a digital version of the work in question.

• Wood, S.L. 1982. The bark and ambrosia beetles of North and Central America (Coleoptera: Scolytidae), a taxonomic monograph. Great Basin Nat. Mem. 6:1-1356. [91]. (data capture complete)
• Wood, S.L., Bright,D.E. 1992. A catalog of Scolytidae and Platypodidae (Coleoptera), Part 2. Taxonomic Index. Great Basin Nat. Mem. 13:1-1553 (vol. A, B). [42]
• Bright, D.E., Skidmore, R.E. 1997. A Catalog of Scolytidae and Platypodidae (Coleoptera), Supplement 1 (1990-1994). NRC Research Press, 368 pp. [10]

The following are references from which host and distribution data have been input into the database. If one of the above monographs or catalogs also appears in this list, it means that most relevant collection event data have been included.

1. . . .
2. Anonymous, 2010. Michigan State University. SCAN: Symbiota Collections of Artropods Network. http://symbiota4.acis.ufl.edu/scan/portal/collections/misc/collprofiles.php?collid=76: -.
3. Atkinson, T.H.; Saucedo C.; E, Martínez F.; E, Burgos S., A 1986. Coleópteros Scolytidae y Platypodidae asociados con las comunidades vegetales de clima templado y frío en el estado de Morelos. Acta Zool. Mex. 17: 1-58.
4. Bright, D.E., Stark, R.W. 1973. The bark and ambrosia beetles of California. Coleoptera: Scolytidae and Platypodidae. Bull. Calif. Insect Surv. 16: 1-169.
5. EDRR 2011. USFS Early Detection Rapid Response Database. http://www.fs.fed.us/invasivespecies/earlydetection.shtml/.
6. EDRR 2014. USFS Early Detection Rapid Response Database. http://www.fs.fed.us/invasivespecies/earlydetection.shtml/.
7. Equihua M., A, Burgos S., A. 1993. Notas sobre la distribución de Scolytidae (Coleoptera) de México y el sur de Texas (EUA). Agrociéncia ser.Proteccion Vegetal 4(3: 305-318.
8. Gast,S.J.; Furniss,M.M.; Johnson,J.B.; Ivie,M.A. 1989. List of Montana Scolytidae (Coleoptera) and notes on new records. Great Basin Nat. 49(3): 381-386.
9. University of Arizona Insect Collection 2016. SCAN: Symbiota Collections of Arthropods Network. http://symbiota4.acis.ufl.edu/scan/portal/collections/misc/collprofiles.php?collid=11.
10. Wood, S.L. 1982. The bark and ambrosia beetles of North and Central America (Coleoptera: Scolytidae), a taxonomic monograph. Great Basin Nat. Mem. 6: 1-1356.
11. Wood, S.L.; Bright, D.E. 1992. A catalog of Scolytidae and Platypodidae (Coleoptera), Part 2. Taxonomic Index (Volumes A,B). Great Basin Nat. Mem. 13: 1-1553.

Maps automatically open at the center of the plotted points and the scale is set to encompass all map points. Maps can be resized (scale bar at upper left) and the center moved (place cursor over map and drag) to see other parts of the distribution of the species. Clicking on a map marker will pull up collection event data and a literature citation if present. If any errors are found, please refer to the "series code" which is a unique identifier for a database record in any communications). Coordinates have not been entered for all collection records. Localities outside the New World are not plotted, even though they are listed in the distribution summary and in the table of records.

No.	Collection data	Publication
North America
1->	, no state / prov. data; Pinaceae: Picea sp.	[unknown (0) (Wood, S.L. 1982: 91)]
2->	, no state / prov. data; Pinaceae: Picea engelmanni	[unknown (0) (Wood, S.L.; Bright, D.E. 1992: 42)]
3->	, no state / prov. data; Pinaceae: Pseudotsuga menziesii	[unknown (0) (Wood, S.L.; Bright, D.E. 1992: 42)]
4->	Canada, British Columbia, Coldwater River	[unknown (1) (Wood, S.L. 1982: 91)]
5->	Canada, British Columbia, Kelowna	[unknown (1) (Wood, S.L. 1982: 91)]
6->	Canada, British Columbia, Midday Valley	[unknown (1) (Wood, S.L. 1982: 91)]
7->	Canada, British Columbia, Midday Valley, Merritt; 10/VI/1925; Stanley, J.; Pinaceae: Pinus ponderosa	[UAIC (4) (University of Arizona Insect Collection 2016: 0)]
8->	Canada, British Columbia, Peachland	[unknown (1) (Wood, S.L. 1982: 91)]
9->	Canada, British Columbia, Summerland	[unknown (1) (Wood, S.L. 1982: 91)]
10->	Canada, British Columbia, Vancouver	[unknown (1) (Wood, S.L. 1982: 91)]
11->	México, no state / prov. data; Pinaceae: Pinus leiophylla	[unknown (0) (Wood, S.L. 1982: 92)]
12->	México, Baja California, Corona Abajo, San Pedro Martir; 18/VII/1963; Sleeper, E.L.	[CAS (16) (TH Atkinson-2007, unpublished)]
13->	México, Baja California, Corona Abajo, San Pedro Martir; 18/VI/1983; Sleeper, E.L.	[CAS (16) (TH Atkinson-2007, unpublished)]
14->	México, Baja California, Sierra de Juárez; 1/XII/1987; Campos B., R.; Pinaceae: Pinus jeffreyi	[unknown () (Equihua M., A, Burgos S., A. 1993: 306)]
15->	México, Chiapas, San Cristobal de las Casas; 16/IV/1984; Saucedo C., E.; Pinaceae: Pinus sp.	[CEAM () (Atkinson, T.H.; Equihua M., A. 1985: 228)]
16->	México, Chiapas, San Cristobal, 5 mi SE Hwy. 190; 25/V/1983; O'Brien, C.W.; O'Brien, L.B.; Marshall, G.B.	[CAS (2) (TH Atkinson-2007, unpublished)]
17->	México, Chiapas, San Cristobal, 5 mi SE, Hwy. 190; 25/V/1983; O'Brien, C.W.; O'Brien, L.B.; Marshall, G.B.; Pinaceae: Pinus sp.	[CAS (2) (TH Atkinson-2007, unpublished)]
18->	México, Chiapas, San Cristobal, 8 mi (13 km) E; 31/VII/1957; Chemsak, J.A.	[unknown (1) (Wood, S.L. 1982: 92)]
19->	México, Chihuahua, Creel (Sierra Madre Mtns. nr. Creel); 3/VII/1972; Giulani, D.	[CAS (2) (TH Atkinson-2007, unpublished)]
20->	México, Chihuahua, La Magdalena; 27/IV/1981; Furniss, M.M.; Cibrían T., D.; Pinaceae: Pinus ayacahuite	[UACC () (Atkinson, T.H.; Equihua M., A. 1988: 92)]
21->	México, Chihuahua, Peña Blanca; 23/IV/1981; Cibrián T., D.; Pinaceae: Pinus chihuahuana	[UACC () (Atkinson, T.H.; Equihua M., A. 1988: 92)]
22->	México, Distrito Federal, Cd. México	[unknown (1) (Wood, S.L. 1982: 92)]
23->	México, Durango, Durango, 60 mi (96 km) W; 5/VI/1965; Wood, S.L.; Pinaceae: Pinus	[USNM (1) (Wood, S.L. 1982: 92)]
24->	México, Durango, El Salto, 1.5 mi W; 19/VIII/1963; Sleeper, E.L.	[CAS (1) (TH Atkinson-2007, unpublished)]
25->	México, Durango, El Salto, 1.5 mi W; 19/VIII/1963; Sleeper, E.L.	[CAS (1) (TH Atkinson-2007, unpublished)]
26->	México, Guerrero, Guerrero (estado)	[Unknown (1) (Wood, S.L. 1982: 92)]
27->	México, Guerrero, Omilteme; Smith, H.H.	[unknown (1) (Wood, S.L. 1982: 92)]
28->	México, Hidalgo, Hidalgo (estado)	[unknown (1) (Wood, S.L.; Bright, D.E. 1992: 42)]
29->	México, México, Carr. México-Puebla, km 56; 28/VI/1961	[CEAM () (Atkinson, T.H.; Equihua M., A. 1985: 69)]
30->	México, México, Carr. México-Puebla, km 56; 15/IV/1961	[CEAM () (Atkinson, T.H.; Equihua M., A. 1985: 69)]
31->	México, México, Carr. México-Puebla, km 57; 8/I/1961; Pinaceae: Pinus sp.	[UACC () (Atkinson, T.H.; Equihua M., A. 1985: 69)]
32->	México, México, Hwy 15, 6 mi E state line with Michoacán; 7/VII/1982; Ivie, M.A.	[TAMU (1) (TH Atkinson-, unpublished)]
33->	México, México, Parque Nal. Zoquiapan; 3/XII/1979; Atkinson, T.H.; Pinaceae: Pinus hartwegii	[CEAM () (Atkinson, T.H.; Equihua M., A. 1985: 69)] [FSCA (2) (Atkinson, T.H.; Equihua M., A. 1985: 69)]
34->	México, México, Parque Nal. Zoquiapan; 4/X/1980; Equihua M., A.; Pinaceae: Pinus montezumae	[CEAM () (Atkinson, T.H.; Equihua M., A. 1985: 69)]
35->	México, México, Parque Nal. Zoquiapan; 3/I/1973; Sánchez, V.	[CEAM () (Atkinson, T.H.; Equihua M., A. 1985: 69)]
36->	México, México, Parque Nal. Zoquiapan; 9/V/1973; Olayo, M.; Pinaceae: Pinus hartwegii	[CEAM () (Atkinson, T.H.; Equihua M., A. 1985: 69)]
37->	México, México, Parque Nal. Zoquiapan; 20/IX/1973; Cibrián T., D.; Pinaceae: Pinus hartwegii	[UACC () (Atkinson, T.H.; Equihua M., A. 1985: 69)]
38->	México, México, Parque Nal. Zoquiapan; 12/IX/1974; Cibrián T., D.; Pinaceae: Pinus hartwegii	[UACC () (Atkinson, T.H.; Equihua M., A. 1985: 69)]
39->	México, México, Real de Arriba; V/1933; Hinton, H.E.; Usinger, R.L.	[BMNH (3) (TH Atkinson-2016, unpublished)]
40->	México, México, Toluca, 31 mi W, Hwy 15; 6/VIII/1982; O'Brien, C.W.; O'Brien, L.B.; Wibmer, G.; Pinaceae: Pinus sp.	[CAS (1) (TH Atkinson-2012, unpublished)]
41->	México, Michoacán, Cheran, 6 mi N; 5/VII/1985; Wooley, J.M.; Zolnerowich, G.	[TAMU (1) (TH Atkinson-2019, unpublished)]
42->	México, Morelos, Chamilpa; Pinaceae: Pinus montezumae	[CEAM () (Atkinson, T.H.; Saucedo C.; E, Martínez F.; E, Burgos S., A 1986: 15)]
43->	México, Morelos, Chamilpa; Pinaceae: Pinus montezumae	[UAEM () (Atkinson, T.H.; Saucedo C.; E, Martínez F.; E, Burgos S., A 1986: 15)]
44->	México, Morelos, Coajomulco	[CEAM () (Atkinson, T.H.; Saucedo C.; E, Martínez F.; E, Burgos S., A 1986: 15)]
45->	México, Morelos, Coajomulco, al norte; Pinaceae: Pinus sp.	[CEAM () (Atkinson, T.H.; Saucedo C.; E, Martínez F.; E, Burgos S., A 1986: 15)]
46->	México, Morelos, Fierro del Toro (al suroeste); Pinaceae: Pinus sp.	[CEAM () (Atkinson, T.H.; Saucedo C.; E, Martínez F.; E, Burgos S., A 1986: 15)]
47->	México, Oaxaca, Miahuatlan, 5 km N; 18/IX/1989; Baranowski, R.M.; Pinaceae: Pinus sp.	[ZMLU (1) (TH Atkinson-2017, unpublished)]
48->	México, Oaxaca, Suchixtepec, 11 mi S, Hwy. 175; 2/VI/1983; O'Brien, C.W.; O'Brien, L.B.; Marshall, G.B.; Pinaceae: Pinus sp.	[FSCA (8) (TH Atkinson-2007, unpublished)]
49->	México, Puebla, Texmelucán, 21 km W; 14/VII/1953; Wood, S.L.; Pinaceae: Pinus	[unknown (1) (Wood, S.L. 1982: 92)]
50->	México, Puebla, Xalitzintla, 3 km W; 26/III/1976; Haswell, R.H.; Pinaceae: Pinus sp.	[TAMU (2) (TH Atkinson-2012, unpublished)]
51->	México, unknown, Mexico (country)	[IRSNB (1: holotype) (Wood, S.L. 1982: 92)]
52->	México, Veracruz, Cofre de Perote; 6/VI/1983; Atkinson, T.H.; Equihua M., A.; Pinaceae: Pinus sp.	[CEAM () (Atkinson, T.H.; Equihua M., A. 1985: 228)] [TAMU (2) (Atkinson, T.H.; Equihua M., A. 1985: 228)]
53->	México, Veracruz, Las Vigas, 10 mi S; 15/VII/1971; Newton, A.; Pinaceae: Pinus sp.	[FMNH (1) (TH Atkinson-2022, unpublished)]
54->	México, Veracruz, Veracruz (estado)	[TAMU (2) (TH Atkinson-, unpublished)]
55->	United States, no state / prov. data; Pinaceae: Pinus ponderosa	[unknown (0) (Wood, S.L. 1982: 92)]
56->	United States, Alaska, Kenai Peninsula	[unknown (1: holotype) (Wood, S.L. 1982: 91)]
57->	United States, Alaska, Kenai Peninsula	[unknown (1: holotype) (Wood, S.L. 1982: 90)]
58->	United States, Arizona, Arizona (state)	[Unknown (1) (Wood, S.L. 1982: 92)]
59->	United States, Arizona, Brady Spring; 15/VII/1980; Olson, C.A.	[UAIC (1) (University of Arizona Insect Collection 2016: 0)]
60->	United States, Arizona, Camp Geronimo; 11/VI/1941; Grassman, P.C.	[UAIC (1) (University of Arizona Insect Collection 2016: 0)]
61->	United States, Arizona, Chiricahua Mtns.	[unknown (1) (Wood, S.L. 1982: 91)]
62->	United States, Arizona, Chiricahua Mtns.; 19/VI/1971; Stephan, K.	[FSCA (2) (TH Atkinson-2013, unpublished)]
63->	United States, Arizona, Chiricahua Mtns.; 4/V/1969; Stephan, K.	[FSCA (1) (TH Atkinson-2013, unpublished)]
64->	United States, Arizona, Chiricahua Mtns., Barfoot Scout Camp; 24/V/1970; Stephan, K.	[FSCA (1) (TH Atkinson-2013, unpublished)]
65->	United States, Arizona, Coconino N.F.	[unknown (1) (Wood, S.L. 1982: 91)]
66->	United States, Arizona, Coronado N.F.	[unknown (1) (Wood, S.L. 1982: 91)]
67->	United States, Arizona, E. Turkey Creek; 16/VI/1978; McCleve, S.; Pinaceae: Pinus sp.	[UAIC (1) (University of Arizona Insect Collection 2016: 0)]
68->	United States, Arizona, East side Marshall Gulch; V/1965; Lindsey, M.L.	[UAIC (14) (University of Arizona Insect Collection 2016: 0)]
69->	United States, Arizona, FS 300, 0.2 mi W Hwy 260; 15/X/2011; Warner, W.B.; Pinaceae: Pinus ponderosa	[WBWC (1) (TH Atkinson-2013, unpublished)]
70->	United States, Arizona, Ft. Valley For. Turp. Area; 5/1959; Nutting, W.L.; Pinaceae: Pinus ponderosa	[UAIC (3) (University of Arizona Insect Collection 2016: 0)]
71->	United States, Arizona, Grand Canyon	[unknown (1) (Wood, S.L. 1982: 91)]
72->	United States, Arizona, Huachuca Mtns.	[unknown (1) (Wood, S.L. 1982: 91)]
73->	United States, Arizona, Kaibab N.F.	[unknown (1) (Wood, S.L. 1982: 91)]
74->	United States, Arizona, Lakeside	[unknown (1) (Wood, S.L. 1982: 91)]
75->	United States, Arizona, Mt. Graham; 18/V/1969; Stephan, K.	[FSCA (1) (TH Atkinson-2013, unpublished)]
76->	United States, Arizona, Mt. Graham; 25/IV/1971; Stephan, K.	[FSCA (1) (TH Atkinson-2013, unpublished)]
77->	United States, Arizona, Payson; 31/V/1972; Stephan, K.	[FSCA (1) (TH Atkinson-2013, unpublished)]
78->	United States, Arizona, Prescott N.F.	[unknown (1) (Wood, S.L. 1982: 91)]
79->	United States, Arizona, Rincon Mtns.; 11/IV/1976; Stephan, K.	[FSCA (1) (TH Atkinson-2013, unpublished)]
80->	United States, Arizona, Rustler's Park; 10/VIII/2019; Schnepp, K.E.; Pinaceae: Pinus sp.	[KESC (1) (TH Atkinson-2020, unpublished)]
81->	United States, Arizona, Rustler's Park; 16/VIII/2019; Schnepp, K.E.; Pinaceae: Pinus sp.	[KESC (1) (TH Atkinson-2020, unpublished)]
82->	United States, Arizona, Rustlers Park; 14/VIII/1970; McCleve, S.; Riley, C.M.; Pinaceae: Pinus sp.	[UAIC (5) (University of Arizona Insect Collection 2016: 0)]
83->	United States, Arizona, Rustlers Park; 8/VI/1974; McCleve, S.; Pinaceae: Pinus sp.	[UAIC (10) (University of Arizona Insect Collection 2016: 0)]
84->	United States, Arizona, Rustlers Park; 14/VIII/1974; McCleve, S.; Pinaceae: Pinus sp.	[UAIC (2) (University of Arizona Insect Collection 2016: 0)]
85->	United States, Arizona, Rustlers Park; 31/VIII/1974; McCleve, S.; Pinaceae: Pinus sp.	[UAIC (17) (University of Arizona Insect Collection 2016: 0)]
86->	United States, Arizona, Rustlers Park; 19/VIII/1973; Nelson, G.H.; Pinaceae: Pinus ponderosa	[FSCA (9) (TH Atkinson-2018, unpublished)]
87->	United States, Arizona, Sta. Catalina Mtns.	[unknown (1) (Wood, S.L. 1982: 91)]
88->	United States, Arizona, Sta. Catalina Mtns.; 7/IX/1968; Stephan, K.	[FSCA (1) (TH Atkinson-2013, unpublished)]
89->	United States, Arizona, Sta. Catalina Mtns.; 20/X/1968; Stephan, K.	[FSCA (1) (TH Atkinson-2013, unpublished)]
90->	United States, Arizona, Sta. Catalina Mtns.; 10/XI/1968; Stephan, K.	[FSCA (3) (TH Atkinson-2013, unpublished)]
91->	United States, Arizona, Sta. Catalina Mtns.; 29/X/1974; Stephan, K.	[FSCA (2) (TH Atkinson-2013, unpublished)]
92->	United States, Arizona, Strayhorse Campground; 30/V/1970; Stephan, K.	[FSCA (1) (TH Atkinson-2013, unpublished)]
93->	United States, Arizona, Strayhorse Campground; 9/VIII/1975; Stephan, K.	[FSCA (1) (TH Atkinson-2013, unpublished)]
94->	United States, Arizona, Williams	[unknown (1) (Wood, S.L. 1982: 91)]
95->	United States, California, Alta	[unknown (1) (Wood, S.L. 1982: 91)]
96->	United States, California, Bass Lake	[unknown (1) (Wood, S.L. 1982: 91)]
97->	United States, California, Big Pines Park	[unknown (1) (Wood, S.L. 1982: 91)]
98->	United States, California, Black Mtn., Lassen N.F.	[unknown (1) (Wood, S.L. 1982: 91)]
99->	United States, California, Blue Lake	[unknown (1) (Wood, S.L. 1982: 91)]
100->	United States, California, Burney	[unknown (1) (Wood, S.L. 1982: 91)]
101->	United States, California, Calaveras Co.	[unknown (1) (Bright, D.E., Stark, R.W. 1973: 22)]
102->	United States, California, Charlton Flat	[unknown (1) (Wood, S.L. 1982: 91)]
103->	United States, California, Charlton Flat; 10/VII/1950; Estes, D.R.	[TAMU (2) (TH Atkinson-1987, unpublished)]
104->	United States, California, Cherry Hill Campground, 5 mi NE Butte Meadows; 24/V/1974; Andrews, F.G.	[CSCA (5) (TH Atkinson-2014, unpublished)]
105->	United States, California, Chester	[unknown (1) (Wood, S.L. 1982: 91)]
106->	United States, California, Chiquito Basin; 1/V/1914; Pinaceae: Pinus ponderosa	[UAIC (20) (University of Arizona Insect Collection 2016: 0)]
107->	United States, California, Chiquito Basin; 1/V/1924	[UAIC (3) (University of Arizona Insect Collection 2016: 0)]
108->	United States, California, Dorrington	[unknown (1) (Wood, S.L. 1982: 91)]
109->	United States, California, Fallen Leaf Lake	[unknown (1) (Wood, S.L. 1982: 91)]
110->	United States, California, Fresno Co.	[unknown (1) (Bright, D.E., Stark, R.W. 1973: 22)]
111->	United States, California, Grass Valley	[unknown (1) (Wood, S.L. 1982: 91)]
112->	United States, California, Groveland; 18/V/2012; Bulaon, B.; trap: Lindgren Funnel ethanol	[unknown (1) (EDRR 2014: 0)]
113->	United States, California, Grover Hot Springs Campground; 11/V/2006; Cline, A.R.; Kerr, P.H.; trap: malaise	[CSCA (1) (TH Atkinson-2014, unpublished)]
114->	United States, California, Hackamore	[unknown (1) (Wood, S.L. 1982: 91)]
115->	United States, California, Jenks Lake; 6/V/2011; Coleman, T.; trap alpha pinene + ethanol	[unknown (1) (EDRR 2011: 0)]
116->	United States, California, Julian; 29/VI/1970; Stephan, K.	[FSCA (1) (TH Atkinson-2013, unpublished)]
117->	United States, California, Kern Co.	[unknown (1) (Bright, D.E., Stark, R.W. 1973: 22)]
118->	United States, California, Kings Beach, Griff Creek; 26/V/1985; Andrews, F.G.; Pinaceae: Pinus jeffreyi	[CSCA (52) (TH Atkinson-2014, unpublished)] [UTIC (8) (TH Atkinson-2014, unpublished)]
119->	United States, California, Lassen N.F.	[unknown (1) (Wood, S.L. 1982: 91)]
120->	United States, California, Mariposa Co.	[unknown (1) (Bright, D.E., Stark, R.W. 1973: 22)]
121->	United States, California, Mather	[unknown (1) (Wood, S.L. 1982: 91)]
122->	United States, California, Mather; 15/VI/1969; Stephan, K.	[FSCA (7) (TH Atkinson-2013, unpublished)]
123->	United States, California, Meadow Valley	[unknown (1) (Wood, S.L. 1982: 91)]
124->	United States, California, Mineral	[unknown (1) (Wood, S.L. 1982: 91)]
125->	United States, California, Mineral King	[unknown (1) (Bright, D.E., Stark, R.W. 1973: 22)]
126->	United States, California, Mineral King; 25/V/1994; Baranowski, R.M.	[ZMLU (2) (TH Atkinson-2017, unpublished)]
127->	United States, California, Mono Co.	[unknown (1) (Bright, D.E., Stark, R.W. 1973: 22)]
128->	United States, California, Oakland	[unknown (1) (Wood, S.L. 1982: 91)]
129->	United States, California, Pickett's Junction, 5mi SE on W Fork Carson Riv.; 10/VII/1985; Hardy, A.R.; Andrews, F.G.	[CSCA (3) (TH Atkinson-2014, unpublished)]
130->	United States, California, Pollock Pines	[unknown (1) (Wood, S.L. 1982: 91)]
131->	United States, California, Sagehen Ck., nr. Hobart Mills; 24/VI/1964; Gillogly, A.R.	[TAMU (1) (TH Atkinson-2003, unpublished)]
132->	United States, California, Sagehen Ck., nr. Hobart Mills; 17/VI/1964; Seminoff, S.G.	[CSCA (1) (TH Atkinson-2014, unpublished)]
133->	United States, California, San Bernardino Co.	[unknown (1) (Bright, D.E., Stark, R.W. 1973: 22)]
134->	United States, California, San Diego	[unknown (1) (Wood, S.L. 1982: 91)]
135->	United States, California, Sequoia N.P.	[unknown (1) (Wood, S.L. 1982: 91)]
136->	United States, California, Shaver Lake Heights; 23/VII/1972; Gilbert, A.J.	[CSCA (1) (TH Atkinson-2014, unpublished)]
137->	United States, California, Siskiyou Co.	[unknown (1) (Bright, D.E., Stark, R.W. 1973: 22)]
138->	United States, California, Susanville; 13/V/2010; Cluck, D.; Garcia, A.; trap alpha pinene + ethanol	[unknown (3) (EDRR 2011: 0)]
139->	United States, California, Susanville; 11/V/2012; Cluck, D.; trap: Lindgren Funnel alpha pinene + ethanol	[unknown (1) (EDRR 2014: 0)]
140->	United States, California, Tahquitz (SanJacinto Mts.)	[unknown (1) (Wood, S.L. 1982: 91)]
141->	United States, California, Trinity Co.	[unknown (1) (Bright, D.E., Stark, R.W. 1973: 22)]
142->	United States, California, Truckee	[unknown (1) (Wood, S.L. 1982: 91)]
143->	United States, California, Walker Mine	[unknown (1) (Wood, S.L. 1982: 91)]
144->	United States, California, Westwood; 26/IV/2007; Smith, S.; Cluck, D.; trap alpha pinene + ethanol	[unknown (14) (EDRR 2011: 0)]
145->	United States, California, Yosemite Valley; 28/V/2014; Bulaon, B.; trap: Lindgren Funnel alpha pinene + ethanol	[unknown (1) (EDRR 2014: 0)]
146->	United States, California, Yuba Co.	[unknown (1) (Bright, D.E., Stark, R.W. 1973: 22)]
147->	United States, Colorado, Allenspark; 14/VI/2011; Daniel, Z.; trap alpha pinene + ethanol	[unknown (1) (EDRR 2011: 0)]
148->	United States, Colorado, Allenspark; 3/V/2012; Daniel, Z.; trap: Lindgren Funnel ethanol	[unknown (1) (EDRR 2014: 0)]
149->	United States, Colorado, Bailey	[unknown (1) (Wood, S.L. 1982: 91)]
150->	United States, Colorado, Blackhawk; 3/V/2012; Daniel, Z.; trap: Lindgren Funnel ethanol	[unknown (1) (EDRR 2014: 0)]
151->	United States, Colorado, Boulder	[unknown (1) (Wood, S.L. 1982: 91)]
152->	United States, Colorado, Boulder; 13/VI/2011; Daniel, Z.; trap alpha pinene + ethanol	[unknown (1) (EDRR 2011: 0)]
153->	United States, Colorado, Caribou; 17/V/2012; Daniel, Z.; trap: Lindgren Funnel ethanol	[unknown (1) (EDRR 2014: 0)]
154->	United States, Colorado, Carter Lake; 4/V/2012; Rowley, T.; trap: Lindgren Funnel alpha pinene + ethanol	[unknown (1) (EDRR 2014: 0)]
155->	United States, Colorado, Estes Park	[unknown (1) (Wood, S.L. 1982: 91)]
156->	United States, Colorado, Estes Park; 13/VI/2011; Rowley, T.; trap alpha pinene + ethanol	[unknown (3) (EDRR 2011: 0)]
157->	United States, Colorado, Estes Park; 4/V/2012; Rowley, T.; trap: Lindgren Funnel ethanol	[unknown (1) (EDRR 2014: 0)]
158->	United States, Colorado, Estes Park; 20/V/2012; Rowley, T.; trap: Lindgren Funnel exotic ips lure	[unknown (1) (EDRR 2014: 0)]
159->	United States, Colorado, Evergreen	[unknown (1) (Wood, S.L. 1982: 91)]
160->	United States, Colorado, Fort Collins; 14/VI/2011; Rowley, T.; trap alpha pinene + ethanol	[unknown (3) (EDRR 2011: 0)]
161->	United States, Colorado, Fort Collins; 4/V/2012; Rowley, T.; trap: Lindgren Funnel alpha pinene + ethanol	[unknown (1) (EDRR 2014: 0)]
162->	United States, Colorado, Fort Collins; 4/V/2012; Rowley, T.; trap: Lindgren Funnel ethanol	[unknown (1) (EDRR 2014: 0)]
163->	United States, Colorado, Gilpin; 16/VI/2011; Daniel, Z.; trap alpha pinene + ethanol	[unknown (6) (EDRR 2011: 0)]
164->	United States, Colorado, Glennhaven	[unknown (1) (Wood, S.L. 1982: 91)]
165->	United States, Colorado, Golden	[unknown (1) (Wood, S.L. 1982: 91)]
166->	United States, Colorado, Livermore; 14/VI/2011; Rowley, T.; trap alpha pinene + ethanol	[unknown (0) (EDRR 2011: 0)]
167->	United States, Colorado, Longdraw; 2/VII/2012; Rowley, T.; trap: Lindgren Funnel alpha pinene + ethanol	[unknown (1) (EDRR 2014: 0)]
168->	United States, Colorado, Longmont; 2/V/2012; Daniel, Z.; trap: Lindgren Funnel ethanol	[unknown (1) (EDRR 2014: 0)]
169->	United States, Colorado, Nederland; 14/VI/2011; Daniel, Z.; trap alpha pinene + ethanol	[unknown (4) (EDRR 2011: 0)]
170->	United States, Colorado, Norwood	[unknown (1) (Wood, S.L. 1982: 91)]
171->	United States, Colorado, Pingree Park	[unknown (1) (Wood, S.L. 1982: 91)]
172->	United States, Colorado, Red Feather Lakes; 14/VI/2011; Rowley, T.; trap alpha pinene + ethanol	[unknown (1) (EDRR 2011: 0)]
173->	United States, Colorado, Stove Praire; 14/VI/2011; Rowley, T.; trap alpha pinene + ethanol	[unknown (1) (EDRR 2011: 0)]
174->	United States, Colorado, Uncompahgre N.F.	[unknown (1) (Wood, S.L. 1982: 91)]
175->	United States, Idaho, Athol; 3/V/1971	[unknown (1) (Wood, S.L. 1982: 91)] [TAMU (7) (Wood, S.L. 1982: 91)]
176->	United States, Idaho, Coeur d'Alene	[unknown (1) (Wood, S.L. 1982: 91)]
177->	United States, Idaho, Harpster Grade	[unknown (1) (Wood, S.L. 1982: 91)]
178->	United States, Idaho, Moscow	[unknown (1) (Wood, S.L. 1982: 91)]
179->	United States, Idaho, Post Falls; 16/IV/2008; Eckberg, T.; trap alpha pinene + ethanol	[unknown (3) (EDRR 2011: 0)]
180->	United States, Idaho, Potlatch	[unknown (1) (Wood, S.L. 1982: 91)]
181->	United States, Idaho, Sand Point; 1/V/2008; Eckberg, T.; trap alpha pinene + ethanol	[unknown (3) (EDRR 2011: 0)]
182->	United States, Idaho, Smith's Ferry	[unknown (1) (Wood, S.L. 1982: 91)]
183->	United States, Idaho, Tamarack	[unknown (1) (Wood, S.L. 1982: 91)]
184->	United States, Idaho, Westmond	[unknown (1) (Wood, S.L. 1982: 91)]
185->	United States, Idaho, Whitebird Pass	[unknown (1) (Wood, S.L. 1982: 91)]
186->	United States, Montana, Cow Creek, Custer N.F.; 1/VI/1988; Furniss, M.M.; Gast, S.J.; Pinaceae: Pinus ponderosa	[WFBM (1) (Gast,S.J.; Furniss,M.M.; Johnson,J.B.; Ivie,M.A. 1989: 384)]
187->	United States, Montana, Hamilton	[unknown (1) (Wood, S.L. 1982: 91)]
188->	United States, Montana, Libby, 3 mi S.; 13/VII/1979; Haswell, R.H.; Pinaceae: Pinus ponderosa	[TAMU (1) (RH Haswell-1979, unpublished)]
189->	United States, Montana, Sula	[unknown (1) (Wood, S.L. 1982: 91)]
190->	United States, Nevada, Reno; 22/IV/2009; Little, R.; trap alpha pinene + ethanol	[unknown (15) (EDRR 2011: 0)]
191->	United States, Nevada, Round Hill; 28/IV/2009; Durham, G.; trap alpha pinene + ethanol	[unknown (10) (EDRR 2011: 0)]
192->	United States, New Mexico, Cloudcroft	[unknown (1) (Wood, S.L. 1982: 91)]
193->	United States, New Mexico, Lincoln N.F.	[unknown (1) (Wood, S.L. 1982: 91)]
194->	United States, New Mexico, Mt. Taylor	[unknown (1) (Wood, S.L. 1982: 91)]
195->	United States, New Mexico, New Mexico (state)	[Unknown (1) (Wood, S.L. 1982: 92)]
196->	United States, New Mexico, Pinos Altos; 4/VII/1975; Stephan, K.	[FSCA (1) (TH Atkinson-2013, unpublished)]
197->	United States, Oregon, Ashland	[unknown (1) (Wood, S.L. 1982: 91)]
198->	United States, Oregon, Bend; 19/IV/2007; Hoffman, J.; trap alpha pinene + ethanol	[unknown (5) (EDRR 2011: 0)]
199->	United States, Oregon, Blue Mts.	[unknown (1) (Wood, S.L. 1982: 91)]
200->	United States, Oregon, Corvallis	[unknown (1) (Wood, S.L. 1982: 91)]
201->	United States, Oregon, Gilchrist; 5/VIII/1997	[MSUC (1) (Anonymous, 2010: 0)]
202->	United States, Oregon, Green Springs	[unknown (1) (Wood, S.L. 1982: 91)]
203->	United States, Oregon, Keno	[unknown (1) (Wood, S.L. 1982: 91)]
204->	United States, Oregon, Madras; 14/IV/2007; Hoffman, J.; trap alpha pinene + ethanol	[unknown (1) (EDRR 2011: 0)]
205->	United States, Oregon, Minam Campground; 12/IX/1974; Haswell, R.H.	[TAMU (8) (TH Atkinson-2012, unpublished)]
206->	United States, Oregon, Pine Cr., Blue Mtns.; 4/VIII/1975; Haswell, R.H.; Pinaceae: Pinus ponderosa	[TAMU (6) (TH Atkinson-2012, unpublished)]
207->	United States, Oregon, Pine Cr., Blue Mtns.; 5/VIII/1974; Haswell, R.H.	[TAMU (6) (TH Atkinson-2012, unpublished)]
208->	United States, Oregon, Salem; 7/IV/2011; Adams, T.; trap alpha pinene + ethanol	[unknown (1) (EDRR 2011: 0)]
209->	United States, Oregon, Santiam Pass	[unknown (1) (Wood, S.L. 1982: 91)]
210->	United States, Oregon, Sisters	[unknown (1) (Wood, S.L. 1982: 91)]
211->	United States, Oregon, Summit Prairie	[unknown (1) (Wood, S.L. 1982: 91)]
212->	United States, Oregon, Warm Springs Indian Reservation, 4 mi N; 3/VII/1970; Turnbow, R.H.; Pinaceae: Pinus ponderosa	[unknown (1) (Wood, S.L. 1982: 91)] [TAMU (2) (Wood, S.L. 1982: 91)]
213->	United States, South Dakota, Black Hills	[unknown (1) (Wood, S.L. 1982: 91)]
214->	United States, South Dakota, Custer	[unknown (1) (Wood, S.L. 1982: 91)]
215->	United States, South Dakota, Deadwood; 29/IV/2009; Halverson; trap alpha pinene + ethanol	[unknown (4) (EDRR 2011: 0)]
216->	United States, South Dakota, Hill City	[unknown (1) (Wood, S.L. 1982: 91)]
217->	United States, South Dakota, Mt. Rushmore; 21/VII/1977; Haswell, R.H.; Pinaceae: Pinus sp.	[TAMU (3) (RH Haswell-1979, unpublished)]
218->	United States, South Dakota, Rapid City; 8/V/2009; Halverson; trap alpha pinene + ethanol	[unknown (1) (EDRR 2011: 0)]
219->	United States, South Dakota, Roubaix; 15/V/2009; Halverson; trap alpha pinene + ethanol	[unknown (1) (EDRR 2011: 0)]
220->	United States, South Dakota, Spearfish	[unknown (1) (Wood, S.L. 1982: 91)]
221->	United States, South Dakota, Sturgis; 29/IV/2009; Halverson; trap alpha pinene + ethanol	[unknown (1) (EDRR 2011: 0)]
222->	United States, South Dakota, Whitewood; 1/V/2009; Halverson; trap alpha pinene + ethanol	[unknown (1) (EDRR 2011: 0)]
223->	United States, Utah, Ashley N.F.	[unknown (1) (Wood, S.L. 1982: 91)]
224->	United States, Utah, Long Hollow, Dixie N.F.	[unknown (1) (Wood, S.L. 1982: 91)]
225->	United States, Washington, Columbia Co.	[unknown (1) (Wood, S.L. 1982: 91)]
226->	United States, Washington, Kooskooskie	[unknown (1) (Wood, S.L. 1982: 91)]
227->	United States, Washington, Kooskooskie; 18/X/1945; Chastain, G.	[UAIC (1) (University of Arizona Insect Collection 2016: 0)]
228->	United States, Washington, Kooskooskie; 24/IV/1949; Chastain, G.; Pinaceae: Pinus ponderosa	[UAIC (2) (University of Arizona Insect Collection 2016: 0)]
229->	United States, Washington, Kooskooskie; 17/V/1940; Ritz, F.	[FSCA (2) (TH Atkinson-2018, unpublished)]
230->	United States, Washington, Naches	[unknown (1) (Wood, S.L. 1982: 91)]
231->	United States, Washington, Preston Creek (Wenatchee N.F.)	[unknown (1) (Wood, S.L. 1982: 91)]
232->	United States, Washington, Roslyn; 13/IV/2007; Fisher; trap alpha pinene + ethanol	[unknown (1) (EDRR 2011: 0)]
233->	United States, Wyoming, Centennial	[unknown (1) (Wood, S.L. 1982: 91)]