Dendrocranulus cucurbitae (LeConte 1879)  


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SYNONYMY


Taxonomic Comments.   Species of this genus are poorly described and diagnostic characters are subtle. Dendrocranulus cucurbitae, D. rudis and D. confinis are very similar and need to be re-examined. Based on specimens originally identified by Wood, many specimens from central Mexico have been placed in this species and need to be re-examined critically.


  • Dendrocranulus cucurbitae (LeConte 1879) Bull. U.S. Geol. Surv. 5: 519
  • Xylocleptes cucurbitae LeConte 1879. Bull. U.S. Geol. Surv. 5: 519
  • Dendrocranulus californicus (Hopkins 1915) U.S. Dep. Agric. Tech. Bull. (99): 44
  • Xylocleptes californicus Hopkins 1915. U.S. Dep. Agric. Tech. Bull. (99): 44
  • Xylocleptes punctatus Hopkins 1915. U.S. Dep. Agric. Tech. Bull. (99): 44
  • Xylocleptes venturina Hopkins 1915. U.S. Dep. Agric. Tech. Bull. (99): 44

DISTRIBUTION.

Numbers in parentheses after each geographic unit are the number of distinct collection events in the database for that unit. For exotic species generally only countries are listed for localities outside the New World. For further information on published sources of distribution, check the REFERENCES section.


Distribution Comments.   Arid and semi-arid regions of the southwestern U.S and Mexico. This genus is in need of revision and it is possible that populations from central Mexico and Central America may be a different species. Commonly found in cultivated species, especially chayote (Sechium edule) and species of Luffa.


  • Central America (2)
    Nicaragua (2): Estelí (2);
  • North America (87)
    (2): no state / prov. data (2); México (38): Baja California (2), Chihuahua (1), México (1), Michoacán (5), Morelos (21), Oaxaca (3), Puebla (1), Sonora (4); United States (47): Arizona (4), California (36), New Mexico (1), Oregon (1), Texas (3), Utah (2);


HOSTS

Numbers in parentheses after each host family, genus, or collection method are the number of distinct collection events in the database for that host or method.


  • No host information (36);
  • Cucurbitaceae (41): (13): no species data (13); Cucurbita (7): argyrosperma (2), digitata (1), foetidissima (3), pepo (1); Cylanthera (1): multiflora (1); Echinocystis (2): macrocarpa (1), sp. (1); Luffa (7): acutangula (3), cylindrica (4); Marah (2): macrocarpa (1), sp. (1); Sechium (6): edule (6); Vaseyanthus (3): insularis (3);
  • litter (2): berlese (2): no species data (1), oak litter (1);
  • trap (10) (4): alpha pinene + ethanol (4); bottle (2): no details (2); Lindgren Funnel (3): ethanol (1), exotic ips lure (2); malaise (1): no details (1);


REFERENCES

The following are important recent monographs, catalogs, and supplements to catalogs that refer to this species. The specific page on which the reference is made is shown in pink at the end of the reference. In the case of Wood (1982) and Wood & Bright (1992) clicking on the reference page will link to a digital version of the work in question.


  • Wood, S.L. 1982. The bark and ambrosia beetles of North and Central America (Coleoptera: Scolytidae), a taxonomic monograph. Great Basin Nat. Mem. 6:1-1356. [718]. (data capture complete)
  • Wood, S.L., Bright,D.E. 1992. A catalog of Scolytidae and Platypodidae (Coleoptera), Part 2. Taxonomic Index. Great Basin Nat. Mem. 13:1-1553 (vol. A, B). [551]

The following are references from which host and distribution data have been input into the database. If one of the above monographs or catalogs also appears in this list, it means that most relevant collection event data have been included.


  1. . . .
  2. Anonymous 2012. Bugguide.net . Bugguide.net (downloaded March, 2012).
  3. Atkinson, T.H. 2019. Escarabajos descortezadores y ambrosiales (Coleoptera: Curculionidae: Scolytinae, Platypodinae) de Sonora, México (Bark and Ambrosia Beetles (Coleoptera: Curculionidae: Scolytinae, Platypodinae) of Sonora, México). Dugesiana 26(1): 41-49.
  4. Atkinson, T.H.; Martínez F., E, Saucedo C., E, Burgos S., A 1986. Scolytidae y Platypodidae (Coleoptera) asociados a selva baja caducifolia y comunidades derivadas en el estado de Morelos. Fol. Entomol. Mex. 69: 41-82.
  5. Bright, D.E., Skidmore, R.E. 2002. A Catalog of Scolytidae and Platypodidae (Coleoptera), Supplement 2 (1995-1999). NRC Research Press. Ottawa, Ontario, Canada.
  6. Bright, D.E., Stark, R.W. 1973. The bark and ambrosia beetles of California. Coleoptera: Scolytidae and Platypodidae. Bull. Calif. Insect Surv. 16: 1-169.
  7. Bright, D.E.; Skidmore, R.S. 1997. A Catalog of Scolytidae and Platypodidae (Coleoptera), Supplement 1 (1990-1994). A Catalog of Scolytidae and Platypodidae (Coleoptera), Supplement 1 (1990-1994): -.
  8. EDRR 2011. USFS Early Detection Rapid Response Database. http://www.fs.fed.us/invasivespecies/earlydetection.shtml/.
  9. EDRR 2014. USFS Early Detection Rapid Response Database. http://www.fs.fed.us/invasivespecies/earlydetection.shtml/.
  10. Equihua M., A, Burgos S., A. 1993. Notas sobre la distribución de Scolytidae (Coleoptera) de México y el sur de Texas (EUA). Agrociéncia ser.Proteccion Vegetal 4(3: 305-318.
  11. Hopkins, A.D. 1915. Classification of the Cryphalinae with descriptions of new genera and species. (99): 1-75.
  12. Lázaro Dzul, M.O.; Equihua Martinez, A.; Romero Nápoles, J.; Gonzalez H., H.; Alvarado, D.; Macías Samaño, J.; Castañeda Vildóla, A.; Atkinson, T,H, 2020. Diversity of Scolytinae (Coleoptera: Curculionidae) at three Sites in the Central Avocado-Producing Region of Michoacán, Mexico. Coleopterists Bull. 74 (2): 454-462.
  13. University of Arizona Insect Collection 2016. SCAN: Symbiota Collections of Arthropods Network. http://symbiota4.acis.ufl.edu/scan/portal/collections/misc/collprofiles.php?collid=11.
  14. Wood, S.L. 1982. The bark and ambrosia beetles of North and Central America (Coleoptera: Scolytidae), a taxonomic monograph. Great Basin Nat. Mem. 6: 1-1356.

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Maps automatically open at the center of the plotted points and the scale is set to encompass all map points. Maps can be resized (scale bar at upper left) and the center moved (place cursor over map and drag) to see other parts of the distribution of the species. Clicking on a map marker will pull up collection event data and a literature citation if present. If any errors are found, please refer to the "series code" which is a unique identifier for a database record in any communications). Coordinates have not been entered for all collection records. Localities outside the New World are not plotted, even though they are listed in the distribution summary and in the table of records.



Show Table   Collection Records Go to images  Go to information summary  Go to distribution map   


No.Collection dataPublication
Central America
1-> Nicaragua, Estelí, La Almaciguera; 13/III/2010; Atkinson, T.H.; Cucurbitaceae: Sechium edule[TAMU (11) (TH Atkinson-2014, unpublished)] [UTIC (11) (TH Atkinson-2014, unpublished)]
2-> Nicaragua, Estelí, La Almaciguera; 14/III/2010; Atkinson, T.H.; Cucurbitaceae[UNAC (21) (TH Atkinson-2014, unpublished)]
North America
3-> , no state / prov. data; Cucurbitaceae: Cucurbita digitata[unknown (0) (Wood, S.L. 1982: 718)]
4-> , no state / prov. data; Cucurbitaceae: Echinocystis macrocarpa[unknown (0) (Wood, S.L. 1982: 718)]
5-> México, Baja California, Isla Sn. Lorenzo Sur; 6/V/1985; Atkinson, T.H.; Cucurbitaceae: Vaseyanthus insularis[CNIN () (Atkinson, T.H.; Equihua M., A. 1988: 88)]
6-> México, Baja California, Isla Sn. Pedro Mártir; 4/V/1985; Atkinson, T.H.; Cucurbitaceae: Vaseyanthus insularis[TAMU (8) (Atkinson, T.H.; Equihua M., A. 1988: 88)] [CNIN (23) (Atkinson, T.H.; Equihua M., A. 1988: 88)]
7-> México, Chihuahua, Meoqui, 100 km NE[unknown (1) (Wood, S.L. 1982: 718)]
8-> México, México, San Miguel Tlaixpan; 4/I/1980; Atkinson, T.H.; Cucurbitaceae: Sechium edule[CEAM (6) (Atkinson, T.H.; Equihua M., A. 1985: 87) (Wood, S.L. 1982: 718) (TH Atkinson-2018, unpublished)]
9-> México, Michoacán, Gabriel Zamora; 1/XI/1980; Atkinson, T.H.; Equihua M., A.; Cucurbitaceae: Luffa acutangula[CEAM () (Atkinson, T.H.; Equihua M., A. 1985: 236) (Wood, S.L. 1982: 718)]
10-> México, Michoacán, Lombardía; 1/XI/1980; Atkinson, T.H.; Equihua M., A.; Cucurbitaceae: Luffa acutangula[CEAM (8) (TH Atkinson-2018, unpublished)]
11-> México, Michoacán, Uruapan; 16/V/1981; Equihua M., A.; Cucurbitaceae: Sechium edule[CEAM (1) (Atkinson, T.H.; Equihua M., A. 1985: 236) (TH Atkinson-2018, unpublished)]
12-> México, Michoacán, Uruápan; VII/2016; Lázaro Dzul, M.O.; trap: bottle[unknown (1) (Lázaro Dzul, M.O.; Equihua Martinez, A.; Romero Nápoles, J.; Gonzalez H., H.; Alvarado, D.; Macías Samaño, J.; Castañeda Vildóla, A.; Atkinson, T,H, 2020: 0)]
13-> México, Michoacán, Ziracuaretiro; VII/2016; Lázaro Dzul, M.O.; trap: bottle[unknown (1) (Lázaro Dzul, M.O.; Equihua Martinez, A.; Romero Nápoles, J.; Gonzalez H., H.; Alvarado, D.; Macías Samaño, J.; Castañeda Vildóla, A.; Atkinson, T,H, 2020: 0)]
14-> México, Morelos, Actopan; 12/X/1983; Saucedo C., E.; Martínez F., E.[CEAM (3) (Atkinson, T.H.; Martínez F., E, Saucedo C., E, Burgos S., A 1986: 60) (TH Atkinson-2018, unpublished)]
15-> México, Morelos, Carr. Xochimilco-Oaxtepec, km 58; 22/I/1980; Atkinson, T.H.; Cucurbitaceae: Cucurbita pepo[CEAM (4) (Atkinson, T.H.; Martínez F., E, Saucedo C., E, Burgos S., A 1986: 60) (TH Atkinson-2018, unpublished)]
16-> México, Morelos, Cerro la Herradura; Burgos S., A.; Cucurbitaceae[UAEM () (Atkinson, T.H.; Martínez F., E, Saucedo C., E, Burgos S., A 1986: 60)]
17-> México, Morelos, Chamilpa; 26/X/1982; Burgos S., A.; Saucedo C., E.[CEAM (1) (Atkinson, T.H.; Martínez F., E, Saucedo C., E, Burgos S., A 1986: 60) (TH Atkinson-2018, unpublished)]
18-> México, Morelos, Chamilpa; Cucurbitaceae[UAEM () (Atkinson, T.H.; Martínez F., E, Saucedo C., E, Burgos S., A 1986: 60)]
19-> México, Morelos, Cuernavaca; 17/IX/1982; Burgos S., A.; Cucurbitaceae: Luffa cylindrica[CEAM (2) (Atkinson, T.H.; Martínez F., E, Saucedo C., E, Burgos S., A 1986: 60) (TH Atkinson-2018, unpublished)]
20-> México, Morelos, Cuernavaca; 15/IX/1982; Burgos S., A.; Cucurbitaceae[CEAM () (Atkinson, T.H.; Martínez F., E, Saucedo C., E, Burgos S., A 1986: 60) (TH Atkinson-2018, unpublished)]
21-> México, Morelos, Cuernavaca; 23/VIII/1982; Burgos S., A.; Cucurbitaceae[CEAM (2) (Atkinson, T.H.; Martínez F., E, Saucedo C., E, Burgos S., A 1986: 60) (TH Atkinson-2018, unpublished)]
22-> México, Morelos, Cuernavaca; 26/XII/1979; Atkinson, T.H.; Cucurbitaceae: Luffa acutangula[CEAM (4) (Atkinson, T.H.; Martínez F., E, Saucedo C., E, Burgos S., A 1986: 60) (Wood, S.L. 1982: 718) (TH Atkinson-2018, unpublished)]
23-> México, Morelos, Cuernavaca; Cucurbitaceae: Luffa cylindrica[CEAM () (Atkinson, T.H.; Martínez F., E, Saucedo C., E, Burgos S., A 1986: 60)]
24-> México, Morelos, Cuernavaca; Burgos S., A.; Cucurbitaceae: Luffa cylindrica[UAEM () (Atkinson, T.H.; Martínez F., E, Saucedo C., E, Burgos S., A 1986: 60)]
25-> México, Morelos, Cuernavaca; 12/X/1982; Burgos S., A.; Cucurbitaceae[CEAM (3) (TH Atkinson-2018, unpublished)]
26-> México, Morelos, Cuernavaca; 7/XI/1982; Burgos S., A.; Saucedo C., E.; Cucurbitaceae[CEAM (2) (TH Atkinson-2018, unpublished)]
27-> México, Morelos, Cuernavaca; 18/III/1982; Burgos S., A.; Cucurbitaceae: Luffa cylindrica[CEAM (2) (TH Atkinson-2018, unpublished)]
28-> México, Morelos, El Regadio; 20/V/1984; Curiel, M.[CEAM (1) (Atkinson, T.H.; Martínez F., E, Saucedo C., E, Burgos S., A 1986: 60) (TH Atkinson-2018, unpublished)]
29-> México, Morelos, Morelos (estado)[FSCA (4) (TH Atkinson-1988, unpublished)]
30-> México, Morelos, Tetela del Volcan; 4/V/1985; Equihua M., A.; Estrada V., E.; Cucurbitaceae[CEAM (6) (TH Atkinson-2018, unpublished)]
31-> México, Morelos, Ticuman; Burgos S., A.; Saucedo C., E.; Martínez F., E.; Cucurbitaceae: Sechium edule[CEAM (2) (Atkinson, T.H.; Martínez F., E, Saucedo C., E, Burgos S., A 1986: 60) (TH Atkinson-2018, unpublished)]
32-> México, Morelos, Tlayacapan; 10/IX/1982; Burgos S., A.; Cucurbitaceae[CEAM (1) (Atkinson, T.H.; Martínez F., E, Saucedo C., E, Burgos S., A 1986: 60) (TH Atkinson-2018, unpublished)]
33-> México, Morelos, Tlayacapan; 19/X/1980; Atkinson, T.H.; Cucurbitaceae: Cylanthera multiflora[CEAM (4) (Atkinson, T.H.; Martínez F., E, Saucedo C., E, Burgos S., A 1986: 60) (TH Atkinson-2018, unpublished)] [FSCA (4) (Atkinson, T.H.; Martínez F., E, Saucedo C., E, Burgos S., A 1986: 60)]
34-> México, Morelos, Tlayacapan; 10/IX/1982; Tejas, A.; Tejas, A.; Cucurbitaceae[CEAM (3) (Atkinson, T.H.; Martínez F., E, Saucedo C., E, Burgos S., A 1986: 60) (TH Atkinson-2018, unpublished)]
35-> México, Oaxaca, Pinotepa, Piedra Blanca; 24/X/1986; Burgos S., A.[CEAM (4) (Equihua M., A, Burgos S., A. 1993: 308) (TH Atkinson-2018, unpublished)]
36-> México, Oaxaca, Sta. Catarina Lachatao; 25/I/2013; Atkinson, T.H.; Cucurbitaceae: Sechium edule[ASUT (12) (TH Atkinson-2014, unpublished)]
37-> México, Oaxaca, Sta. Ma. Chimalapa, BEnito Juárez; 6/II/1984; Equihua M., A.; Cucurbitaceae: Sechium edule[TAMU (16) (TH Atkinson-2010, unpublished)] [CEAM (1) (TH Atkinson-2018, unpublished)]
38-> México, Puebla, Palmar del Bravo; 18/I/2013; Atkinson, T.H.; Cucurbitaceae[UTIC (8) (TH Atkinson-2014, unpublished)]
39-> México, Sonora, Alamos Hwy, Aduana turnoff; 31/I/2016; Atkinson, T.H.; Smith, S.M.; Cucurbitaceae: Cucurbita argyrosperma[UTIC (6) (Atkinson, T.H. 2019: 45)] [UAIC (8) (Atkinson, T.H. 2019: 45)] [BMNH (12) (Atkinson, T.H. 2019: 45)]
40-> México, Sonora, Isla San Esteban; 5/V/1985; Atkinson, T.H.; Cucurbitaceae: Vaseyanthus insularis[CNIN (23) (Atkinson, T.H.; Equihua M., A. 1988: 88) (Atkinson, T.H. 2019: 45)]
41-> México, Sonora, Isla Tiburón; 11/II/1986[CNIN (5) (TH Atkinson-2018, unpublished)]
42-> México, Sonora, Tecoripa, 10 km W; 10/III/2016; Atkinson, T.H.; Cucurbitaceae: Cucurbita argyrosperma[UTIC (6) (TH Atkinson-2016, unpublished)] [CEAM (4) (TH Atkinson-2016, unpublished)] [UAIC (4) (TH Atkinson-2016, unpublished)]
43-> United States, Arizona, Arizona (state)[MCZ (7) (TH Atkinson-1988, unpublished)]
44-> United States, Arizona, Bonita[unknown (1) (Wood, S.L. 1982: 718)]
45-> United States, Arizona, Hwy 191, 0.5 mi w jct Dragoon Rd.; 10/V/2002; Turnbow, R.H.; Cucurbitaceae: Cucurbita foetidissima[FSCA (8) (TH Atkinson-2007, unpublished)]
46-> United States, Arizona, Hwy 191, 0.5 mi w jct Dragoon Rd.[FSCA (1) (MC Thomas-2012, downloaded from Bugguide)]
47-> United States, California, Aguanga; 3/XI/1991; Valcarce, V.; Hawks, D.A.; Kirkendall, L.A.; Cucurbitaceae: Marah macrocarpa[TAMU (27) (TH Atkinson-1992, unpublished)]
48-> United States, California, Alum Rock Pk.[CWOB (1) (Bright, D.E., Stark, R.W. 1973: 74)]
49-> United States, California, Andrew Molera St. Pk.; 27/VI/2006; Cline, A.R.; Hauser, M.; trap: malaise[THAC (6) (TH Atkinson-2014, unpublished)] [CSCA (10) (TH Atkinson-2014, unpublished)]
50-> United States, California, Azusa[MCZ (1) (TH Atkinson-1988, unpublished)]
51-> United States, California, Azusa[unknown (1) (Wood, S.L. 1982: 718)] [OSU (1) (Bright, D.E., Stark, R.W. 1973: 74)]
52-> United States, California, Azusa; 27/X/1934; McClay, A.T.[UAIC (8) (University of Arizona Insect Collection 2016: 0)]
53-> United States, California, Bautista Canyon; 3/II/1980; Cooper, K.W.; litter: berlese[CSCA (1) (TH Atkinson-2014, unpublished)]
54-> United States, California, Big Sur, 4 mi S; 23/VI/1990; Haswell, R.H.[TAMU (1) (TH Atkinson-2012, unpublished)]
55-> United States, California, Darrah, 2 mi NE; 17/XI/1975; Gilbert, A.J.; litter: berlese oak litter[CSCA (1) (TH Atkinson-2014, unpublished)]
56-> United States, California, Desert Center[unknown (1) (Wood, S.L. 1982: 718)]
57-> United States, California, Desert Center[SJSC (1) (Bright, D.E., Stark, R.W. 1973: 74)]
58-> United States, California, El Cajon; 5/VI/2014; Coleman, T.W.; trap: Lindgren Funnel exotic ips lure[unknown (1) (EDRR 2014: 0)]
59-> United States, California, Glendale; 13/V/2000; Wellso, S.G.[TAMU (1) (TH Atkinson-2010, unpublished)]
60-> United States, California, Glendale[UCDC (1) (Bright, D.E., Stark, R.W. 1973: 74)]
61-> United States, California, Greenhorn Summit, 8.2 mi W; 2/X/2004; Turnbow, R.H.; Cucurbitaceae: Marah sp.[FSCA (25) (TH Atkinson-2007, unpublished)] [RHTC (4) (TH Atkinson-2008, unpublished)]
62-> United States, California, Lagunas[unknown (1) (Wood, S.L. 1982: 718)]
63-> United States, California, Martinez[unknown (1) (Wood, S.L. 1982: 718)]
64-> United States, California, Martinez; Cucurbitaceae[CAS (1) (Bright, D.E., Stark, R.W. 1973: 74)]
65-> United States, California, Newton[unknown (1) (Wood, S.L. 1982: 718)]
66-> United States, California, North Claremont; 17/V/2010; Coleman, T.; trap alpha pinene + ethanol[unknown (2) (EDRR 2011: 0)]
67-> United States, California, Oakland Hills[CAS (1) (Bright, D.E., Stark, R.W. 1973: 74)]
68-> United States, California, Palo Alto, Stanford University[SJSC (1) (Bright, D.E., Stark, R.W. 1973: 74)]
69-> United States, California, Pasadena[MCZ (1) (TH Atkinson-1988, unpublished)]
70-> United States, California, Pasadena[unknown (1) (Wood, S.L. 1982: 718)] [CAS (1) (Bright, D.E., Stark, R.W. 1973: 74)]
71-> United States, California, Playa del Rey[unknown (1) (Wood, S.L. 1982: 718)]
72-> United States, California, Playa del Rey; Cucurbitaceae: Cucurbita foetidissima[OSU (1) (Bright, D.E., Stark, R.W. 1973: 74)]
73-> United States, California, Pomona[MCZ (1) (TH Atkinson-1988, unpublished)]
74-> United States, California, Pomona; 1900; Hubbard, H.G.; Schwarz, E.A.[USNM (1: holotype, female) (Wood, S.L. 1982: 718) (Hopkins, A.D. 1915: 44) (TH Atkinson-2017, unpublished)]
75-> United States, California, Rancho Penasquitos; 17/V/2010; Coleman, T.; trap alpha pinene + ethanol[unknown (1) (EDRR 2011: 0)]
76-> United States, California, San Bernardino; 19/IV/2012; Coleman, T.W.; trap: Lindgren Funnel exotic ips lure[unknown (1) (EDRR 2014: 0)]
77-> United States, California, San Bernardino; 24/V/2013; Coleman, T.W.; trap: Lindgren Funnel ethanol[unknown (1) (EDRR 2014: 0)]
78-> United States, California, Sierra Madre[unknown (1) (Wood, S.L. 1982: 718)]
79-> United States, California, Soledad; 17/III/1977; Oliver, A.; Cucurbitaceae: Echinocystis sp.[CSCA (6) (TH Atkinson-2014, unpublished)]
80-> United States, California, Sorrento Valley; 17/V/2010; Coleman, T.; trap alpha pinene + ethanol[unknown (1) (EDRR 2011: 0)]
81-> United States, California, Trabuco Canyon; 13/V/1972; Cicero, J.N.[FSCA (1) (TH Atkinson-2007, unpublished)]
82-> United States, California, Ventura Co; 1/VI/1904; Hopkins, A.D.; Cucurbitaceae[USNM (1: holotype, female) (Wood, S.L. 1982: 718) (Hopkins, A.D. 1915: 44) (TH Atkinson-2017, unpublished)]
83-> United States, New Mexico, Mesilla; 18/III/; Cockerell, T.D.A.; Cucurbitaceae: Cucurbita foetidissima[USNM (1: holotype, female) (Wood, S.L. 1982: 718) (Hopkins, A.D. 1915: 44) (TH Atkinson-2017, unpublished)]
84-> United States, Oregon, Brookings; 9/II/2011; TMJ; trap alpha pinene + ethanol[unknown (2) (EDRR 2011: 0)]
85-> United States, Texas, Ft. Davis, 8.6 mi W. hwy 118; 7/VIII/1992; Godwin, W.; Riley, E.G.[TAMU (1) (TH Atkinson-2003, unpublished)]
86-> United States, Texas, Kent, 30 km SE ; 22/VII/1989; Anderson, R.[TAMU (1) (TH Atkinson-2003, unpublished)]
87-> United States, Texas, Madera Canyon, 23 mi N Ft. Davis; 19/VII/1974; Bright, D.E.[CNC (18) (Bright, D.E.; Skidmore, R.S. 1997: 120) (Bright, D.E., Skidmore, R.E. 2002: 512)]
88-> United States, Utah, St. George[unknown (1) (Wood, S.L. 1982: 718)]
89-> United States, Utah, Utah (state)[MCZ (1: holotype, female) (Wood, S.L. 1982: 718)]

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