Bark and Ambrosia Beetles of

SYNONYMY

• Ips latidens (LeConte 1874) Trans. Amer. Ent. Soc. 5: 72
• Orthotomicus latidens (LeConte 1874) Trans. Amer. Ent. Soc. 5: 72
• Tomicus latidens LeConte 1874. Trans. Amer. Ent. Soc. 5: 72
• Ips longidens Swaine 1911. Can. Entomol. 43: 214
• Ips guildi Blackman 1922. NY State Coll. For. Tech. Pub. 16: 137

DISTRIBUTION.

Numbers in parentheses after each geographic unit are the number of distinct collection events in the database for that unit. For exotic species generally only countries are listed for localities outside the New World. For further information on published sources of distribution, check the REFERENCES section.

Distribution Comments.   Western North America south to Baja California and Chihuahua; across central Canada to northeastern North America

• North America (198)
  (8): no state / prov. data (8); Canada (9): Alberta (1), British Columbia (1), New Brunswick (3), Nova Scotia (1), Ontario (1), Quebec (1), Saskatchewan (1); México (3): Baja California (1), Chihuahua (2); United States (178): Arizona (4), California (57), Colorado (20), Idaho (6), Maine (1), Massachusetts (2), Michigan (7), Montana (4), Nebraska (1), Nevada (3), New Hampshire (1), New Mexico (8), New York (6), Oregon (14), Pennsylvania (3), South Dakota (1), Utah (31), Washington (6), West Virginia (1), Wyoming (2);

HOSTS

Numbers in parentheses after each host family, genus, or collection method are the number of distinct collection events in the database for that host or method.

• No host information (71);
• Pinaceae (15): Larix (1): laricina (1); Pinus (14): albicaulis (1), contorta (2), edulis (1), flexilis (1), halapensis (1), jeffreyi (1), lambertaina (1), lambertiana (1), monticola (1), ponderosa (3), strobus (1);
• trap (112) (78): alpha pinene + ethanol (78); car net (1): no details (1); light (1): ultraviolet (1); Lindgren Funnel (32): no details (2), alpha pinene + ethanol (9), ethanol (5), exotic ips lure (15), ips-lure (1);

REFERENCES

The following are important recent monographs, catalogs, and supplements to catalogs that refer to this species. The specific page on which the reference is made is shown in pink at the end of the reference. In the case of Wood (1982) and Wood & Bright (1992) clicking on the reference page will link to a digital version of the work in question.

• Wood, S.L. 1982. The bark and ambrosia beetles of North and Central America (Coleoptera: Scolytidae), a taxonomic monograph. Great Basin Nat. Mem. 6:1-1356. [676]. (data capture complete)
• Wood, S.L., Bright,D.E. 1992. A catalog of Scolytidae and Platypodidae (Coleoptera), Part 2. Taxonomic Index. Great Basin Nat. Mem. 13:1-1553 (vol. A, B). [510]

The following are references from which host and distribution data have been input into the database. If one of the above monographs or catalogs also appears in this list, it means that most relevant collection event data have been included.

1. Anonymous, 2010. Michigan State University. SCAN: Symbiota Collections of Artropods Network. http://symbiota4.acis.ufl.edu/scan/portal/collections/misc/collprofiles.php?collid=76: -.
2. Bright, D.E., Stark, R.W. 1973. The bark and ambrosia beetles of California. Coleoptera: Scolytidae and Platypodidae. Bull. Calif. Insect Surv. 16: 1-169.
3. Cognato, A.I.; Barc, N.; Philip, M.; Mech, R.; Smith, A.D.; Galbraith, E.; Storer, A.J.; Kirkendall, L.R. 2009. The native and introduced bark and ambrosia beetles of Michigan (Curculionidae: Scolytinae). Great Lakes Entom.42(3-4): 101-120.
4. Cognato, A.I.; Rogers, S.O.; Teale, S.A. 1995. Species diagnosis and phylogeny of the Ips grandicollis group (Coleoptera: Scolytidae) using random amplified polymorphic DNA. Ann. Entomol. Soc. Am. 88 (4): 397-405.
5. Cognato, A.I.; Sperling, F.A.H. 2000. Phylogeny of Ips DeGeer species (Coleoptera: Scolytidae) inferred from mitochondrial cytochrome oxidase I DNA sequence. Mol. Phylogenet. Evol. 14 (3): 445-460.
6. EDRR 2011. USFS Early Detection Rapid Response Database. http://www.fs.fed.us/invasivespecies/earlydetection.shtml/.
7. EDRR 2014. USFS Early Detection Rapid Response Database. http://www.fs.fed.us/invasivespecies/earlydetection.shtml/.
8. University of Arizona Insect Collection 2016. SCAN: Symbiota Collections of Arthropods Network. http://symbiota4.acis.ufl.edu/scan/portal/collections/misc/collprofiles.php?collid=11.
9. Webster, R., Anderson, R.S., Sweeney, J., DeMerchant, I 2012. New Coleoptera records from New Brunswick, Canada: Anthribidae, Brentidae, Dryophthoridae, Brachyceridae, and Curculionidae, with additions to the fauna of Quebec, Nova Scotia and Prince Edward Island. Zookeys 179: 349-406.
10. Wood, S.L. 1951. The Scolytidae of the Logan Canyon area in Utah and their host plants. Proc.Utah Acad.Sci.Arts Letters 26: 127-128.
11. Wood, S.L. 1982. The bark and ambrosia beetles of North and Central America (Coleoptera: Scolytidae), a taxonomic monograph. Great Basin Nat. Mem. 6: 1-1356.

Maps automatically open at the center of the plotted points and the scale is set to encompass all map points. Maps can be resized (scale bar at upper left) and the center moved (place cursor over map and drag) to see other parts of the distribution of the species. Clicking on a map marker will pull up collection event data and a literature citation if present. If any errors are found, please refer to the "series code" which is a unique identifier for a database record in any communications). Coordinates have not been entered for all collection records. Localities outside the New World are not plotted, even though they are listed in the distribution summary and in the table of records.

No.	Collection data	Publication
North America
1->	, no state / prov. data	[TAMU () (TH Atkinson-, unpublished)]
2->	, no state / prov. data; Pinaceae: Pinus albicaulis	[unknown (0) (Wood, S.L. 1982: 677)]
3->	, no state / prov. data; Pinaceae: Pinus edulis	[unknown (0) (Wood, S.L. 1982: 677)]
4->	, no state / prov. data; Pinaceae: Pinus jeffreyi	[unknown (0) (Wood, S.L. 1982: 677)]
5->	, no state / prov. data; Pinaceae: Pinus lambertiana	[unknown (0) (Wood, S.L. 1982: 677)]
6->	, no state / prov. data; Pinaceae: Pinus monticola	[unknown (0) (Wood, S.L. 1982: 677)]
7->	, no state / prov. data; Pinaceae: Pinus ponderosa	[unknown (0) (Wood, S.L. 1982: 677)]
8->	, no state / prov. data; Pinaceae: Pinus strobus	[unknown (0) (Wood, S.L. 1982: 677)]
9->	Canada, Alberta, Alberta (province)	[unknown (1) (Wood, S.L. 1982: 676)]
10->	Canada, British Columbia, British Columbia (province)	[unknown (1) (Wood, S.L. 1982: 676)]
11->	Canada, New Brunswick, Acadia Res. Forest; 2/VI/2009; Webster, R.P.; Giguère, M.-A.; trap: Lindgren Funnel	[RWC (1) (Webster, R., Anderson, R.S., Sweeney, J., DeMerchant, I 2012: 397)]
12->	Canada, New Brunswick, Fredericton; 8/VI/1925; Simpson, L.J.	[AFC (5) (Webster, R., Anderson, R.S., Sweeney, J., DeMerchant, I 2012: 397)]
13->	Canada, New Brunswick, Tracy, 15 km W Rt. 645; 19/V/2009; Webster, R.P.; Giguère, M.-A.; trap: Lindgren Funnel	[RWC (1) (Webster, R., Anderson, R.S., Sweeney, J., DeMerchant, I 2012: 397)]
14->	Canada, Nova Scotia, Halifax; 16/VI/1999; Smith, G.; trap: Lindgren Funnel ips-lure	[AFC (1) (Webster, R., Anderson, R.S., Sweeney, J., DeMerchant, I 2012: 397)]
15->	Canada, Ontario, Ontario (province)	[unknown (1) (Wood, S.L. 1982: 676)]
16->	Canada, Quebec, Quebec (province)	[unknown (1) (Wood, S.L. 1982: 676)]
17->	Canada, Saskatchewan, Saskatchewan (province)	[unknown (1) (Wood, S.L. 1982: 676)]
18->	México, Baja California, San Pedro Mártir NP; 8/XI/2015; Atkinson, T.H.; Pinaceae: Pinus contorta	[UTIC (2) (TH Atkinson-2016, unpublished)]
19->	México, Chihuahua, Cusárare	[unknown (1) (Wood, S.L. 1982: 676)]
20->	México, Chihuahua, San Juanito	[unknown (1) (Wood, S.L. 1982: 676)]
21->	United States, Arizona, Arizona (state)	[unknown (1) (Wood, S.L. 1982: 676)]
22->	United States, Arizona, Flagstaff; 23/IV/2009; DeGomez, T.; trap alpha pinene + ethanol	[unknown (25) (EDRR 2011: 0)]
23->	United States, Arizona, Tucson; 7/II/1981; Werner, F.G.; Pinaceae: Pinus halapensis	[UAIC (4) (University of Arizona Insect Collection 2016: 0)]
24->	United States, Arizona, White Mtns.; VI/; Duncan, D.K.	[UAIC (1) (University of Arizona Insect Collection 2016: 0)]
25->	United States, California, Anderson; 18/IV/2007; Seybold, S.J.; Owen, D.; trap alpha pinene + ethanol	[unknown (1) (EDRR 2011: 0)]
26->	United States, California, Anderson; 15/VIII/2014; Owen, D.; trap: Lindgren Funnel exotic ips lure	[unknown (1) (EDRR 2014: 0)]
27->	United States, California, Anderson; 14/VI/2013; Owen, D.; trap: Lindgren Funnel exotic ips lure	[unknown (1) (EDRR 2014: 0)]
28->	United States, California, Big Bear; 6/V/2011; Coleman, T.; trap alpha pinene + ethanol	[unknown (1) (EDRR 2011: 0)]
29->	United States, California, Butte Co.	[unknown (1) (Bright, D.E., Stark, R.W. 1973: 84)]
30->	United States, California, Calaveras Co.	[unknown (1) (Bright, D.E., Stark, R.W. 1973: 84)]
31->	United States, California, California (state)	[MCZ (1: holotype, female) (Wood, S.L. 1982: 676)]
32->	United States, California, Camino, Winkelmans Reservoir; 7/III/1965; Seminoff, S.G.	[CSCA (10) (TH Atkinson-2014, unpublished)] [UTIC (4) (TH Atkinson-2014, unpublished)] [UTIC (4) (TH Atkinson-2014, unpublished)] [UTIC (2) (TH Atkinson-2014, unpublished)] [UTIC (2) (TH Atkinson-2014, unpublished)]
33->	United States, California, Camp Connell, 1.8 mi NE; 24/V/1983; Andrews, F.G.; Hardy, A.R.	[CSCA (1) (TH Atkinson-2014, unpublished)]
34->	United States, California, Cherry Hill Campground, 5 mi NE Butte Meadows; 24/V/1974; Andrews, F.G.	[CSCA (5) (TH Atkinson-2014, unpublished)]
35->	United States, California, China Flat Campground; 21/VI/1975; Hardy, W.; Hardy, A.R.	[CSCA (4) (TH Atkinson-2014, unpublished)]
36->	United States, California, Crestline; 3/V/2012; Coleman, T.W.; trap: Lindgren Funnel exotic ips lure	[unknown (1) (EDRR 2014: 0)]
37->	United States, California, Descanso; 10/VII/2014; Coleman, T.W.; trap: Lindgren Funnel exotic ips lure	[unknown (1) (EDRR 2014: 0)]
38->	United States, California, El Dorado Co.	[unknown (1) (Bright, D.E., Stark, R.W. 1973: 84)]
39->	United States, California, Fish Camp; 15/IV/1995; Cognato, A.I.; Pinaceae: Pinus lambertaina	[EMEC (1) (Cognato, A.I.; Sperling, F.A.H. 2000: 457)]
40->	United States, California, Folsom Lake; 11/IV/1965; Seminoff, S.G.	[CSCA (6) (TH Atkinson-2014, unpublished)]
41->	United States, California, Fresno Co.	[unknown (1) (Bright, D.E., Stark, R.W. 1973: 84)]
42->	United States, California, Glenn Co.	[unknown (1) (Bright, D.E., Stark, R.W. 1973: 84)]
43->	United States, California, Groveland; 18/V/2012; Bulaon, B.; trap: Lindgren Funnel exotic ips lure	[unknown (1) (EDRR 2014: 0)]
44->	United States, California, Humboldt Co.	[unknown (1) (Bright, D.E., Stark, R.W. 1973: 84)]
45->	United States, California, Inyo Co.	[unknown (1) (Bright, D.E., Stark, R.W. 1973: 84)]
46->	United States, California, Jenks Lake; 6/V/2011; Coleman, T.; trap alpha pinene + ethanol	[unknown (5) (EDRR 2011: 0)]
47->	United States, California, Kern Co.	[unknown (1) (Bright, D.E., Stark, R.W. 1973: 84)]
48->	United States, California, Lake Co.	[unknown (1) (Bright, D.E., Stark, R.W. 1973: 84)]
49->	United States, California, Madera Co.	[unknown (1) (Bright, D.E., Stark, R.W. 1973: 84)]
50->	United States, California, Mammoth Lakes; 6/VI/2012; Bulaon, B.; trap: Lindgren Funnel exotic ips lure	[unknown (1) (EDRR 2014: 0)]
51->	United States, California, Mariposa Co.	[unknown (1) (Bright, D.E., Stark, R.W. 1973: 84)]
52->	United States, California, Meeks Bay; VI/1977; Andrews, F.G.	[CSCA (1) (TH Atkinson-2014, unpublished)]
53->	United States, California, Modoc Co.	[unknown (1) (Bright, D.E., Stark, R.W. 1973: 84)]
54->	United States, California, Mono Co.	[unknown (1) (Bright, D.E., Stark, R.W. 1973: 84)]
55->	United States, California, Monterey Co.	[unknown (1) (Bright, D.E., Stark, R.W. 1973: 84)]
56->	United States, California, Pacific House, 0.7 mi E; 2/VII/1987; Andrews, F.G.	[CSCA (1) (TH Atkinson-2014, unpublished)]
57->	United States, California, Placer Co.	[unknown (1) (Bright, D.E., Stark, R.W. 1973: 84)]
58->	United States, California, Plumas Co.	[unknown (1) (Bright, D.E., Stark, R.W. 1973: 84)]
59->	United States, California, Plumas-Eureka State Park; 3/VI/1978; Mitchell, B.; Rohde	[UAIC (1) (University of Arizona Insect Collection 2016: 0)]
60->	United States, California, Redding; 24/IV/2008; Synder, C.; Owen, D.; trap alpha pinene + ethanol	[unknown (1) (EDRR 2011: 0)]
61->	United States, California, Redding; 17/V/2013; Snyder, C.; trap: Lindgren Funnel alpha pinene + ethanol	[unknown (1) (EDRR 2014: 0)]
62->	United States, California, Redding; 22/VI/2012; Snyder, C.; trap: Lindgren Funnel alpha pinene + ethanol	[unknown (1) (EDRR 2014: 0)]
63->	United States, California, Riverside Co.	[unknown (1) (Bright, D.E., Stark, R.W. 1973: 84)]
64->	United States, California, San Diego Co.	[unknown (1) (Bright, D.E., Stark, R.W. 1973: 84)]
65->	United States, California, San Luis Obispo	[unknown (1) (Bright, D.E., Stark, R.W. 1973: 84)]
66->	United States, California, Santa Clara Co.	[unknown (1) (Bright, D.E., Stark, R.W. 1973: 84)]
67->	United States, California, Santa Cruz Co.	[unknown (1) (Bright, D.E., Stark, R.W. 1973: 84)]
68->	United States, California, Siskiyou Co.	[unknown (1) (Bright, D.E., Stark, R.W. 1973: 84)]
69->	United States, California, Skyforest; 5/V/2008; Coleman, T.; trap alpha pinene + ethanol	[unknown (1) (EDRR 2011: 0)]
70->	United States, California, Sonora Pass; 30/IV/1987; Andrews, F.G.; Eichlin, T.	[CSCA (1) (TH Atkinson-2014, unpublished)]
71->	United States, California, Susanville; 24/IV/2008; Cluck, D.; trap alpha pinene + ethanol	[unknown (1) (EDRR 2011: 0)]
72->	United States, California, Susanville; 24/V/2012; Cluck, D.; trap: Lindgren Funnel exotic ips lure	[unknown (1) (EDRR 2014: 0)]
73->	United States, California, Susanville; 23/V/2013; Cluck, D.; trap: Lindgren Funnel exotic ips lure	[unknown (1) (EDRR 2014: 0)]
74->	United States, California, Susanville; 6/VI/2014; Cluck, D.; trap: Lindgren Funnel exotic ips lure	[unknown (1) (EDRR 2014: 0)]
75->	United States, California, Trinity Co.	[unknown (1) (Bright, D.E., Stark, R.W. 1973: 84)]
76->	United States, California, Tulare Co.	[unknown (1) (Bright, D.E., Stark, R.W. 1973: 84)]
77->	United States, California, Tuolumne Co.	[unknown (1) (Bright, D.E., Stark, R.W. 1973: 84)]
78->	United States, California, Van Nuys; 17/IV/2007; Seybold, S.J.; trap alpha pinene + ethanol	[unknown (4) (EDRR 2011: 0)]
79->	United States, California, Ventura Co	[unknown (1) (Bright, D.E., Stark, R.W. 1973: 84)]
80->	United States, California, Westwood; 26/IV/2007; Smith, S.; Cluck, D.; trap alpha pinene + ethanol	[unknown (15) (EDRR 2011: 0)]
81->	United States, California, Yosemite NP; 28/V/2014; Bulaon, B.; trap: Lindgren Funnel exotic ips lure	[unknown (1) (EDRR 2014: 0)]
82->	United States, Colorado, Allenspark; 14/VI/2011; Daniel, Z.; trap alpha pinene + ethanol	[unknown (3) (EDRR 2011: 0)]
83->	United States, Colorado, Boulder; 13/VI/2011; Daniel, Z.; trap alpha pinene + ethanol	[unknown (2) (EDRR 2011: 0)]
84->	United States, Colorado, Carter Lake; 4/V/2012; Rowley, T.; trap: Lindgren Funnel alpha pinene + ethanol	[unknown (1) (EDRR 2014: 0)]
85->	United States, Colorado, Colorado (state)	[unknown (1) (Wood, S.L. 1982: 676)]
86->	United States, Colorado, Estes Park; 13/VI/2011; Rowley, T.; trap alpha pinene + ethanol	[unknown (1) (EDRR 2011: 0)]
87->	United States, Colorado, Estes Park; 3/VI/2012; Rowley, T.; trap: Lindgren Funnel ethanol	[unknown (1) (EDRR 2014: 0)]
88->	United States, Colorado, Estes Park; 3/VIII/2012; Rowley, T.; trap: Lindgren Funnel exotic ips lure	[unknown (1) (EDRR 2014: 0)]
89->	United States, Colorado, Fort Collins; 14/VI/2011; Rowley, T.; trap alpha pinene + ethanol	[unknown (1) (EDRR 2011: 0)]
90->	United States, Colorado, Fort Collins; 4/V/2012; Rowley, T.; trap: Lindgren Funnel alpha pinene + ethanol	[unknown (1) (EDRR 2014: 0)]
91->	United States, Colorado, Fort Collins; 4/V/2012; Rowley, T.; trap: Lindgren Funnel ethanol	[unknown (1) (EDRR 2014: 0)]
92->	United States, Colorado, Franktown; 11/IV/2007; Aguayo, I.; trap alpha pinene + ethanol	[unknown (2) (EDRR 2011: 0)]
93->	United States, Colorado, Grand Junction; 23/IV/2007; Aguayo, I.; trap alpha pinene + ethanol	[unknown (2) (EDRR 2011: 0)]
94->	United States, Colorado, Grand Lake; 4/III/1921; Guild, E.R.	[USNM (1: holotype, female) (Wood, S.L. 1982: 676) (TH Atkinson-2018, unpublished)]
95->	United States, Colorado, Longmont; 2/V/2012; Daniel, Z.; trap: Lindgren Funnel ethanol	[unknown (1) (EDRR 2014: 0)]
96->	United States, Colorado, Loveland; 13/VI/2011; Rowley, T.; trap alpha pinene + ethanol	[unknown (3) (EDRR 2011: 0)]
97->	United States, Colorado, Montrose; 1/VI/2007; Hohenstein, J.; trap alpha pinene + ethanol	[unknown (2) (EDRR 2011: 0)]
98->	United States, Colorado, Nederland; 14/VI/2011; Daniel, Z.; trap alpha pinene + ethanol	[unknown (1) (EDRR 2011: 0)]
99->	United States, Colorado, Red Feather Lakes; 14/VI/2011; Rowley, T.; trap alpha pinene + ethanol	[unknown (1) (EDRR 2011: 0)]
100->	United States, Colorado, Stove Praire; 14/VI/2011; Rowley, T.; trap alpha pinene + ethanol	[unknown (2) (EDRR 2011: 0)]
101->	United States, Colorado, Stove Praire; 21/V/2012; Rowley, T.; trap: Lindgren Funnel alpha pinene + ethanol	[unknown (1) (EDRR 2014: 0)]
102->	United States, Idaho, Boise; 6/VI/2008; Schlenker, L.; trap alpha pinene + ethanol	[unknown (1) (EDRR 2011: 0)]
103->	United States, Idaho, East Port; 1/V/2008; Eckberg, T.; trap alpha pinene + ethanol	[unknown (2) (EDRR 2011: 0)]
104->	United States, Idaho, Idaho (state)	[unknown (1) (Wood, S.L. 1982: 676)]
105->	United States, Idaho, Moscow; 15/IV/2008; Looney, D.; trap alpha pinene + ethanol	[unknown (9) (EDRR 2011: 0)]
106->	United States, Idaho, Post Falls; 16/IV/2008; Eckberg, T.; trap alpha pinene + ethanol	[unknown (1) (EDRR 2011: 0)]
107->	United States, Idaho, Twin Falls; 2/VI/2008; Dersch, L.; trap alpha pinene + ethanol	[unknown (1) (EDRR 2011: 0)]
108->	United States, Maine, Bath; 26/III/2009; Urquhart, W.; trap alpha pinene + ethanol	[unknown (1) (EDRR 2011: 0)]
109->	United States, Massachusetts, 04/14/2008; 14/IV/2008; Gooch, K.; trap alpha pinene + ethanol	[unknown (1) (EDRR 2011: 0)]
110->	United States, Massachusetts, Massachussetts (state)	[unknown (1) (Wood, S.L. 1982: 676)]
111->	United States, Michigan, Deward; 3/XII/1917; Pettit; Pinaceae: Larix laricina	[MSUC (1) (Cognato, A.I.; Barc, N.; Philip, M.; Mech, R.; Smith, A.D.; Galbraith, E.; Storer, A.J.; Kirkendall, L.R. 2009: 1) (Anonymous, 2010: 0)]
112->	United States, Michigan, Muskegon St. Park; 5/I/2008; Barc, N.; Smith, A.	[MSUC (1) (Anonymous, 2010: 0)]
113->	United States, Michigan, Muskegon St. Park; 30/V/2008; Barc, N.; Smith, A.	[MSUC (1) (Anonymous, 2010: 0)]
114->	United States, Michigan, Refuge rd.; 6/IV/2008	[MSUC (1) (Anonymous, 2010: 0)]
115->	United States, Michigan, Richfield; 5/V/2008; Mech, R.; trap alpha pinene + ethanol	[unknown (1) (EDRR 2011: 0)]
116->	United States, Michigan, Richfield County Park; 27/V/2008; Mech, R.; trap: Lindgren Funnel alpha pinene + ethanol	[MSUC (1) (Cognato, A.I.; Barc, N.; Philip, M.; Mech, R.; Smith, A.D.; Galbraith, E.; Storer, A.J.; Kirkendall, L.R. 2009: 1) (Anonymous, 2010: 0)]
117->	United States, Michigan, Roscommon; 16/V/2011; Radtke; trap alpha pinene + ethanol	[unknown (1) (EDRR 2011: 0)]
118->	United States, Montana, Billings; 14/IV/2009; Toone, C.; trap alpha pinene + ethanol	[unknown (1) (EDRR 2011: 0)]
119->	United States, Montana, Bozeman; 14/IV/2009; Toone, C.; trap alpha pinene + ethanol	[unknown (1) (EDRR 2011: 0)]
120->	United States, Montana, Great Falls; 15/IV/2009; Toone, C.; trap alpha pinene + ethanol	[unknown (2) (EDRR 2011: 0)]
121->	United States, Montana, Montana (state)	[unknown (1) (Wood, S.L. 1982: 676)]
122->	United States, Nebraska, Chappell; 9/IV/2008; Harrell, M.; trap alpha pinene + ethanol	[unknown (1) (EDRR 2011: 0)]
123->	United States, Nevada, Elko; 28/IV/2009; Little, R.; trap alpha pinene + ethanol	[unknown (3) (EDRR 2011: 0)]
124->	United States, Nevada, Reno; 22/IV/2009; Little, R.; trap alpha pinene + ethanol	[unknown (4) (EDRR 2011: 0)]
125->	United States, Nevada, Round Hill; 28/IV/2009; Durham, G.; trap alpha pinene + ethanol	[unknown (4) (EDRR 2011: 0)]
126->	United States, New Hampshire, New Hampshire (state)	[unknown (1) (Wood, S.L. 1982: 676)]
127->	United States, New Mexico, Albuquerque; 6/V/2008; Sandoval, S.; trap alpha pinene + ethanol	[unknown (1) (EDRR 2011: 0)]
128->	United States, New Mexico, Cloudcroft, 8.05 km E, Rt. 82 ; 11/V/1994; Pinaceae: Pinus ponderosa	[unknown (1) (Cognato, A.I.; Rogers, S.O.; Teale, S.A. 1995: 398)]
129->	United States, New Mexico, Dixon; 8/V/2008; Sandoval, S.; trap alpha pinene + ethanol	[unknown (1) (EDRR 2011: 0)]
130->	United States, New Mexico, Jemez Springs, 6 mi S; 22/VIII/1968; Andrews, F.G.; Noonan, G.R.; trap: light ultraviolet	[CSCA (1) (TH Atkinson-2014, unpublished)]
131->	United States, New Mexico, Los Alamos; 7/V/2008; Sandoval, S.; trap alpha pinene + ethanol	[unknown (1) (EDRR 2011: 0)]
132->	United States, New Mexico, New Mexico (state)	[unknown (1) (Wood, S.L. 1982: 676)]
133->	United States, New Mexico, Ruidoso; 5/V/2008; Sandoval, S.; trap alpha pinene + ethanol	[unknown (3) (EDRR 2011: 0)]
134->	United States, New Mexico, Santa Fe; 9/V/2008; Sandoval, S.; trap alpha pinene + ethanol	[unknown (1) (EDRR 2011: 0)]
135->	United States, New York, Boonville; 5/VI/2014; Cole, R.; Cancelliere, J.; trap: Lindgren Funnel exotic ips lure	[unknown (1) (EDRR 2014: 0)]
136->	United States, New York, Ithaca	[CNC (1: lectotype, male) (Wood, S.L. 1982: 676)]
137->	United States, New York, Mount Morris; 16/V/2007; FH field crew; trap alpha pinene + ethanol	[unknown (1) (EDRR 2011: 0)]
138->	United States, New York, Pound Ridge; 24/V/2012; Denham, J.; trap: Lindgren Funnel ethanol	[unknown (1) (EDRR 2014: 0)]
139->	United States, New York, Rotterdam; 11/IV/2011; Cancelliere, J.; trap alpha pinene + ethanol	[unknown (1) (EDRR 2011: 0)]
140->	United States, New York, Westchester; 16/V/2007; Carnes, H.; trap alpha pinene + ethanol	[unknown (1) (EDRR 2011: 0)]
141->	United States, Oregon, Bend; 19/IV/2007; Hoffman, J.; trap alpha pinene + ethanol	[unknown (1) (EDRR 2011: 0)]
142->	United States, Oregon, Central Point; 12/IX/1997	[MSUC (1) (Anonymous, 2010: 0)]
143->	United States, Oregon, Central Point; 29/VI/1997	[MSUC (1) (Anonymous, 2010: 0)]
144->	United States, Oregon, Gilchrist; 5/VIII/1997	[MSUC (1) (Anonymous, 2010: 0)]
145->	United States, Oregon, Gilchrist; 28/VI/1997	[MSUC (1) (Anonymous, 2010: 0)]
146->	United States, Oregon, Madras; 14/IV/2007; Hoffman, J.; trap alpha pinene + ethanol	[unknown (2) (EDRR 2011: 0)]
147->	United States, Oregon, Ontario; 30/IV/2007; Rogg, H.; trap alpha pinene + ethanol	[unknown (9) (EDRR 2011: 0)]
148->	United States, Oregon, Oregon (state)	[unknown (1) (Wood, S.L. 1982: 676)]
149->	United States, Oregon, Portland; 7/IV/2011; Larsen, K.A.; trap alpha pinene + ethanol	[unknown (1) (EDRR 2011: 0)]
150->	United States, Oregon, Rogue River; 8/IV/2011; Stoner, D.C.; trap alpha pinene + ethanol	[unknown (3) (EDRR 2011: 0)]
151->	United States, Oregon, Salem; 7/IV/2011; Adams, T.; trap alpha pinene + ethanol	[unknown (3) (EDRR 2011: 0)]
152->	United States, Oregon, The Dalles; 13/VI/1997	[MSUC (1) (Anonymous, 2010: 0)]
153->	United States, Oregon, The Dalles; 5/VIII/1997	[MSUC (1) (Anonymous, 2010: 0)]
154->	United States, Oregon, The Dalles; 14/VIII/1997	[MSUC (1) (Anonymous, 2010: 0)]
155->	United States, Pennsylvania, Liberty; 30/V/2013; Laubscher, B.; trap: Lindgren Funnel alpha pinene + ethanol	[unknown (1) (EDRR 2014: 0)]
156->	United States, Pennsylvania, Pennsylvania (state)	[unknown (1) (Wood, S.L. 1982: 676)]
157->	United States, Pennsylvania, Ridley Creek SP ; 30/III/2006; Spichiger, S.; trap alpha pinene + ethanol	[unknown (1) (EDRR 2011: 0)]
158->	United States, South Dakota, Ditch Creek Road and Deerfield Lake, Black Hills National Fo; 28/VI/2015; Schulke, M.; trap: car net	[ZMHB (6) (TH Atkinson-2019, unpublished)]
159->	United States, Utah, Bountiful; 19/V/2011; Grolley, D.; trap alpha pinene + ethanol	[unknown (1) (EDRR 2011: 0)]
160->	United States, Utah, Brigham City; 9/IV/2007; Burfitt; trap alpha pinene + ethanol	[unknown (1) (EDRR 2011: 0)]
161->	United States, Utah, Cleveland; 23/VII/2007; Burfitt; trap alpha pinene + ethanol	[unknown (1) (EDRR 2011: 0)]
162->	United States, Utah, Cottonwood Heights; 16/VII/2007; Burfitt; trap alpha pinene + ethanol	[unknown (35) (EDRR 2011: 0)]
163->	United States, Utah, Danieles; 19/V/2011; Grolley, D.; trap alpha pinene + ethanol	[unknown (1) (EDRR 2011: 0)]
164->	United States, Utah, Dutch John; 8/VIII/2007; Burfitt; trap alpha pinene + ethanol	[unknown (4) (EDRR 2011: 0)]
165->	United States, Utah, Fairview; 15/V/2007; Burfitt; trap alpha pinene + ethanol	[unknown (141) (EDRR 2011: 0)]
166->	United States, Utah, Farmington; 15/III/2007; Burfitt; trap alpha pinene + ethanol	[unknown (7) (EDRR 2011: 0)]
167->	United States, Utah, Farr West; 12/III/2007; Burfitt; trap alpha pinene + ethanol	[unknown (6) (EDRR 2011: 0)]
168->	United States, Utah, Gunnison; 27/VI/2007; Burfitt; trap alpha pinene + ethanol	[unknown (1) (EDRR 2011: 0)]
169->	United States, Utah, Heber; 9/IV/2007; Burfitt; trap alpha pinene + ethanol	[unknown (21) (EDRR 2011: 0)]
170->	United States, Utah, Helper; 23/VII/2007; Burfitt; trap alpha pinene + ethanol	[unknown (1) (EDRR 2011: 0)]
171->	United States, Utah, Hyde Park; 9/IV/2007; Burfitt; trap alpha pinene + ethanol	[unknown (1) (EDRR 2011: 0)]
172->	United States, Utah, Kamas; 27/VII/2007; Burfitt; trap alpha pinene + ethanol	[unknown (1) (EDRR 2011: 0)]
173->	United States, Utah, Layton; 3/VIII/2007; Burfitt; trap alpha pinene + ethanol	[unknown (1) (EDRR 2011: 0)]
174->	United States, Utah, Logan Canyon; 1946; Wood, S.L.; Pinaceae: Pinus contorta	[USNM (1) (Wood, S.L. 1982: 676) (Wood, S.L. 1951: 128)]
175->	United States, Utah, Logan Canyon; 1946; Wood, S.L.; Pinaceae: Pinus flexilis	[USNM (1) (Wood, S.L. 1951: 128)]
176->	United States, Utah, Millcreek; 22/V/2012; Crowther, N.; trap: Lindgren Funnel alpha pinene + ethanol	[unknown (1) (EDRR 2014: 0)]
177->	United States, Utah, Murray; 15/V/2012; Crowther, N.; trap: Lindgren Funnel exotic ips lure	[unknown (1) (EDRR 2014: 0)]
178->	United States, Utah, North Ogden; 23/V/2011; Grolley, D.; trap alpha pinene + ethanol	[unknown (1) (EDRR 2011: 0)]
179->	United States, Utah, Ogden; 19/V/2011; Grolley, D.; trap alpha pinene + ethanol	[unknown (1) (EDRR 2011: 0)]
180->	United States, Utah, Provo; 10/IV/2007; Burfitt; trap alpha pinene + ethanol	[unknown (2) (EDRR 2011: 0)]
181->	United States, Utah, Provo Canyon; 1/VI/2011; Crowther, N.; trap alpha pinene + ethanol	[unknown (1) (EDRR 2011: 0)]
182->	United States, Utah, Salt Lake; 23/III/2007; Burfitt; trap alpha pinene + ethanol	[unknown (3) (EDRR 2011: 0)]
183->	United States, Utah, Salt Lake City; 13/III/2007; Burfitt; trap alpha pinene + ethanol	[unknown (2) (EDRR 2011: 0)]
184->	United States, Utah, Salt Lake City; 7/V/2012; Crowther, N.; trap: Lindgren Funnel ethanol	[unknown (1) (EDRR 2014: 0)]
185->	United States, Utah, South Salt Lake City; 7/V/2012; Crowther, N.; trap: Lindgren Funnel alpha pinene + ethanol	[unknown (1) (EDRR 2014: 0)]
186->	United States, Utah, Washington Terrace; 26/VII/2007; Burfitt; trap alpha pinene + ethanol	[unknown (11) (EDRR 2011: 0)]
187->	United States, Utah, West Valley City; 8/VII/2007; Burfitt; trap alpha pinene + ethanol	[unknown (22) (EDRR 2011: 0)]
188->	United States, Utah, West Valley City; 15/V/2012; Crowther, N.; trap: Lindgren Funnel exotic ips lure	[unknown (1) (EDRR 2014: 0)]
189->	United States, Utah, West Valley City; 15/V/2012; Crowther, N.; trap: Lindgren Funnel exotic ips lure	[unknown (1) (EDRR 2014: 0)]
190->	United States, Washington, Kooskooskie; 17/V/1949; Chastain, C.	[UAIC (4) (University of Arizona Insect Collection 2016: 0)]
191->	United States, Washington, Roslyn; 13/IV/2007; Fisher; trap alpha pinene + ethanol	[unknown (1) (EDRR 2011: 0)]
192->	United States, Washington, Roslyn; 30/VI/1997; Cognato, A.I.; Pinaceae: Pinus ponderosa	[EMEC (1) (Cognato, A.I.; Sperling, F.A.H. 2000: 457)]
193->	United States, Washington, Seattle; 9/IV/2007; Fisher; trap alpha pinene + ethanol	[unknown (1) (EDRR 2011: 0)]
194->	United States, Washington, Terrace Heights; 13/IV/2007; Fisher; trap alpha pinene + ethanol	[unknown (1) (EDRR 2011: 0)]
195->	United States, Washington, Washington (state)	[unknown (1) (Wood, S.L. 1982: 676)]
196->	United States, West Virginia, West Virginia (state)	[unknown (1) (Wood, S.L. 1982: 676)]
197->	United States, Wyoming, Cheyenne; 20/V/2009; Koch, L.; trap alpha pinene + ethanol	[unknown (5) (EDRR 2011: 0)]
198->	United States, Wyoming, Wyoming (state)	[unknown (1) (Wood, S.L. 1982: 676)]